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SMITTISONIAN INSTITUTION.
UNIGED STATES NATIONAL MUSEUM.
PROCEEDINGS
ELLED
alse (5° WF ee
| ‘
OF THE
UNITED STATES NATIONS li TARESEUM. vie \ineus tS
cs
Vo laime «x OS LIE,
PUBLISHED UNDER THE DIRECTION OF THE SMITHSONIAN INSTITUTION.
WASHINGTON: GOVERNMENT PRINTING OFFICE. LOO Lt.
ADVERTISEMENT.
The publications of the National Museum consist of two series: Proceedings and Bulletins.
The Proceedings, the first volume of which was issued in 1878, are intended primarily as a medium of publication for newly-acquired facts in biology, anthropology, and geology, descriptions of new forms of animals and plants acquired by the National Museum, discussions of nomenclature, etc. A volume is issued annually for distribution to libraries, while in view of the importance to science of the prompt publication of descriptions of new species, a limited edition of each paper is printed in pamphlet form in advance.
The present volume is the twenty-third of the series.
The Bulletin, publication of which was begun in 1875, is a series of elaborate papers, issued separately and based for the most part upon collections in the National Museum. They are monographic in scope, and are devoted principally to the discussion of large. zoological groups, bibliographies of eminent -naturalists, reports of expedi- tions, ete.
A quarto form of the Bulletin, known as the ‘‘Special Bulletin,” has been adopted in a few instances in which a larger page was deemed indispensable. |
The Annual Report of the National Museum (being the second vol- ume of the Smithsonian Report) contains papers chiefly of an ethno- logical character, describing collections in the National Museum.
Papers intended for publication by the National Museum are usually referred to an advisory committee, composed as follows: Frederick W. True (chairman), William H. Holmes, George P. Merrill, James E. Benedict, Otis T. Mason, Leonhard Stejneger, Lester F. Ward, and Marcus Benjamin (editor).
S. P. LANGLEY, Secretary of the Smithsonian Institution.
Ill
(tho
TABLE OF CONTENTS.
ApBpott, JAMES Francis. List of Fishes collected in the River Pei-Ho, at Tien-Tsin, China, by Noah Fields Drake, with descriptions of seven new species.—No. 1221. Febru- Rime Se eee hee Rene eh Sere ee Ne is ct a Sela ai
New genus: Culticula.
New species: Toxabramis argentifer, Culticula emmelas, Pseudogobio drakei, Leuciscus sctistius, Parapelecus macherius, Culter tientsin- ensis, Salana hyalocranius.
ASHMEAD, WriittAm H. Classification of the Ichneumon Flies, or the Superfamily Ichneumonoidea.—No. 1206. (JSG) STE SES IG (00 aie a et ER a et 2
Banks, NatHan. Some Spiders and other Arachnida from Southern Arizona.—No. 1223. February 25, 1901? New species: Prosthesima arizonensis, Prosthesima schwarzi, Chemmis unicolor, Dipena parvula, Philodromus marginellus, Chelifer hub-
bardi, Chelanops arizonensis, Thargalia luctuosa.
Benepict, JAMES E. Four new symmetrical Hermit Crabs (Pagurids) from the West India region.—No. 1236. July 1. 1901"
New species: Cancellus ornatus, Cancellus spongicola, Pylocheles parti- tus, Mixtopagurus gilli.
The Hermit Crabs of the Pagurus Bernhardus Type.—No.: 1216... January 19, 1901*
Buscx, Aveust. New species of Moths of the Superfamily Tineina from Florida.—No. 1208. October 15, 19001
New genera: Gnorimoschema, Homaledra, Podiasa, Metriochroa, Chilo- campyla.
New species: Aristotelia ive, Aproaerema crotolariella, Gnorimoschema terracottella, Nealyda pisonix, Nealyda kinzelella, Anacampsis lagun- culariella, Anacampsis argyrothamniella, Trichotaphe melantherella, Trichotaphe condaliavorella, Depressaria amyrisella, Blastobasis gui- landinze, Cosmopteryx ipomoex, Cosmopteryx nigrapunctella, Antispila eugeniella, Homaledra heptathalama, Nepticula condatiafoliella, Nepticula myricafoliella, Leucoptera erythrinella, Leucoptera guettar- della, Podiasa chiococcella, Coptodisca condalix, Bucculatrix ivella, Bedellia minor, Metriochroa psychotriella, Marmara guilandinella, Lithocolletis verbesinella, Coriscium randiella, Chilocampyla dyariella, Eucosmophora sideroxylonella, Gracilaria burserella, Gracilaria sebastianiella, Phyllocnistis intermediella.
CockErRELL, T. D. A. Ona Slug of the Genus Veronicella irom wahitt.—No. 1288. August. 15,1900... ..:..-.-
New species: Veronicella agassizi.
‘Date of publication.
Page.
483-491
o81—
590
TU1-T78
451-466
225-254
855—
536
yar TABLE OF CONTENTS.
Page. CoquiLLerr, D. W. A systematic arrangement of the Fami- lies of the Diptera.—No. 1227. May 2,1901* _..-..----- 653-658
——. New Diptera in the U. S. National Museum.—No. 205.6 ° March 27, 51901 Soe = 33, eee hee ee 593-618
New genera: Pycnoglossa, Plethochzta.
New species: Bolitophila montana, Macrocera nebulosa, Ceroplatus clausus, Platyura inops, Tetragoneura bicolor, Tetragoneura pimpla, Sciophila subcerulea, Sciophila flavohirta, Polylepta tibialis, Docosialongicornis, Docosia obscura, Docosia vittata, Leptomorphus parvulus, Leptomor- phus hyalinus, Acnemia flaveola, Exechia analis, Dynatosoma thora- cica, Chironomus brachialis, Chironomus teniapennis, Chironomus nitidulus, Orthocladius par, Euryenemus scitulus, Tanypus johnsoni, Tanypus bifasciatus, Empis tridentata, Rhamphomyia clauda, Rham- phomyia diversa, Criorhina kincaidi, Scatophaga nubifera, Seatophaga crinita, Hexamitocera flavida, Pycnoglossa flavipennis, Plethochxta varicolor, Anorostoma opaca, Tetanocera setosa, Sciomyza guttata, Sciomyza albovaria, Neuroctena fumida, Dryomyza aristalis, Psila frontalis, Loxocera fumipennis, Phortica vittata.
Dati, Witt1aAM Hratey. Synopsis of the family Cardiide and of the North American species.—No. 1214. January De AGLI. hp ee Ae Ee a ee ae en 381-392
Synopsis of the family Tellinide and of the North American species.—No. 1210. November 14, 1900°_----- 285-326 New species: Tellina( Eurytellina?) georgiana, Fellina (Lnotellina) iher-~ ingi, Tellina (Elliptotellina) americana, Tellina (Angulus) promera, Tellina (Angulus) flagellum, Tellina (Angulus) colorata, Tellina (Angulus) texana, Macoma (Macoma) phenax, Macoma (Psamma- coma) extenuata, Macoma (Psammacoma) tageliformis, Tellina (Merisca) reclusa, Tellina (Elliptotellina) pacifica, Tellina ( Phyl- lodina) pristiphora, Tellina (Eurytellina) leucogonia, Tellina (Moerella) meropsis, Tellina (Moerella) amianta, Tellina (Moerella) paziana, Tellina (Angulus) macneilii, Tellina (Angulus) suffusa, Tellina (Angulus) cerrosiana, Tellina (Angulus) panamensis, Tellina (Angulus) recurva, Tellina (Peronidia) santarose, Macoma krausei, Macoma sitkana, Macoma alaskana, Macoma_ (Psammacoma) panamensis. New names: Tellina (Angulus) carpenteri, Tellina (Oudardia) buttoni.
Synopsis of the Lucinacea and of the American species.—No; 1237. . August 22; 190). sae eee T79-833
New species: Thyasira excavata, Thyasira tomeana, Thyasira magel- lanica, Axinopsis viridis, Diplodonta (Torelli) aleutica, Codakia colpoica, Codakia cubana, Codakia (Jagonia) portoricana, Codakia (Jagonia) galapagana, Codakia (Jagonia) chiquita, Phacoides ( Callucina) bermudensis, Phacoides (Parvilucina) crenella, Phacoides (Bellucina) amiantus, Phacoides (Cavilucina) lamprus, Phacoides (Lucinoma) heroicus, Phacoides ( Parvilucina) approximatus.
New names: Codakia (Jagonia) mexicana, Divaricella perparvula.
1 Date of publication.
TABLE OF CONTENTS. VII
*2 . . . a : Page. Dyar, Harrison G. Life Histories of some North American evans —No: 1209. October 15, 19004. =... 22... _ 955-284 Pr Git, THEODORE. The proper names of Bdellostoma or Hep- rare. No. 1234.8 June 6, 19017 8 2: 2S le 735-738
“JORDAN, David STARR, AND Epwin CHAPIN STARKS. On the relationship of the Lutianoid Fish, Aphareus Furecatus.— Siete Neto etOOT et wee Aa a Fe 719-723
JORDAN, Davip STARR, AND JOHN OTTERBEIN SNYDER. <A list of Fishes collected in Japan by Keinosuke Otaki, and by the United States Steamer Albatross, with descriptions of Fourteen New species.—No. 1213. December 10,19001_ 335-380 New genera: Otakia, Ischikauia, Konosirus, Bryttosus, Eteliscus, Insidi- ator, Trifissus. New species: Chimera phantasma, Gobio biwx, Gobio mayedx, Otakia rasborina, Congrellus meeki, Pseudotolithus mitsukurii, Sebastodes hakodatis, Sebastodes scythropus, Scorpena onaria, Callionymus beni- teguri, Trifissus ioturus, Blennius yatebei, Celorhynchus kishinouyei, Verasper otakii. . List of Fishes collected in 1883 and 1885 by Pierre Louis Jouy and preserved in the United States National Museum, with descriptions of six new species.—No. 1235. STOUT hy TSU AEE Be ca ei ae AI ee ec 139-769
New genera: Chasmias, Watasea. New species: Leuciscus jouyi, Apogon. unicolor, Pomacentrus rathbuni, Aboma tsushime, Chasmias misakius, Watasea sivicola. JORDAN, DaviD STARR, AND JOHN OTTERBEIN SNYDER. <A Review of the Lancelets, Hag-fishes, and Lampreys of
Japan, with a description of two new species.—No. 1233. ramet BR OUNCE e Erie tiesto ee Slee 725-734
New species: Branchiostoma nakagawex, Myxine garmani.
——. A review of the Apodal Fishes or Eels of Japan, with descriptions of nineteen new species.—No. 1239. Pea ES EPA aeE Ul Se OAS ele owes tse ae Je lad 2s 837-890
New genera: Chlevastes, Xyrias, -Emasia.
New species: Synaphobranchus iraconis, Synaphobranchus jenkinsi, Lep- tocephalus erebennus, Leptocephalus kiusiuanus, Leptocephalus riukiu- anus, Leptocephalus nystromi, Leptocephalus retrotinctus, Chlopsis Jierasfer, Murexnichthys owstoni, Murenichthys hatte, Murenichthys aoki, Sphagebranchus moseri, Pisoodonophis zophistius, Nyrias revul- sus, Microdonophis erabo, Ophichthus asakuse, Ophichthus tsuchide, Aimasia lichenosa, Echidna kishinouyei, Uropterygius okinawe.
. Areview of the Cardinal Fishes of Japan. No.1240. error etam ed iis estes ep LI Ue TY 891-913
New genus: Telescopias. New species: Apogon dederleini, Apogon kiensis, Telescopias gilberti.
1 Date of publication.
$A ALLL LLL LLL LAL
a
VALET TABLE OF CONTENTS.
Page. Lucas, Freperic A. A new Dinosaur, Stegosaurus marshi, from the Lower Cretaceous of South Dakota.—No. 1224. Rebruary 26, 19012. <sest o2 ae es 591-592
A New Fossil Cyprinoid, Leuciscus turneri, from the Miocene of Nevada.—No. 1212. November 8, 19001. 833-334
Anew Rhinoceros, Trigonias osborni, from the Mio- cene of South Dakota.—No. 1207. October 9, 1900?____. 221-2294
The Pelvic Girdle of Zeuglodon, Basilosaurus cetoi- des (Owen), with notes on other portions of the skeleton.— Noe iit <November-83*lO00 ta. ac eo ae eee eee 327-831
Lyon Jr., Marcus W. <A comparison of the Osteology of the Jerboas and Jumping Mice.—No. 1228. May 2, 1901'_ 659-668
MecNEILL, JEROME. Revision of the Orthopteran Genus Tri- merotropis:—No:1215.) January (921901 =o eee 398-449
Newspecies: Trimerotropis hyalina, Trimerotropiscristata, Trimerotropis porrecta, Trimerotropiscoquilletti, Trimerotropis tesselata, Trimerotropis calignosa, Trimerotropis albescens, Trimerotropis ferruginea, Trimero- tropis bruneri, Trimerotropis fascicula, Trimerotropis preclara, Tri- merotropis melanoptera, Trimerotropis strenua, Trimerotropis agrestis, Trimerotropis collaris, Trimerotropis fratercula, Trimerotropis saxa- tilis, Trimerotropis pilosa, Trimerotropis nubila, Trimerotropis con- spersa, Trimerotropis vuriegata.
Maxon, WititiawR. A listof Ferns and Fern allies of North America North of Mexico, with principal synonyms and distribution-—No. 1226.+-,May 424.900 *, tu 024.2 eee 619-651
Netson, Exitas. <A Revision of Certain Species of Plants of the Genus Antennaria.—No. 1230. June 4, 1901?______- 697-713
Ricuarpson, Harrier. Key to the Isopods of the Atlantic Coast of North America with Descriptions of New and Little known ‘species.—No. 1222. February 28, 1901'_______- 493-579
New genus: Synuropus.
New name: Frichsonella.
New species: Calathura crenulata, Cirolana obtruncata, Cirolana albida, Corallana sexticornis, Egathoa linguifrons, Spheroma yucatanum, Dynamene angulata, Cilicea linguicauda, Erichsonella floridana, Arc- turus caribbeus, Eurycope caribbea, Synuropus granulatus, Philoscia richmondi, Sphxroniscus portoricensis.
STEJNEGER, Leonnarp. A New Systematic name for the Yel- low Boa of Jamaica.—No. 1218. January 19, 19011. ____. 469-470
New species: Epicrates subflavus.
1 Date of publication.
TABLE OF CONTENTS. Ix cae: ; Page. STEINEGER, LEONHARD. Description of new species of Snake from Clarion Island, West Coast of Mexico.—No. 1231. erm ern Ne gen se ery ee Se T1B-T1T
New species: Bascanion anthonyi.
Diagnosis of a New species of Iguanoid Lizard from Green Cay, Bahama Islands.—No. 1219. January 19, 19017 471 New species: Leiocephalus virescens. . On a new species of Spiny-tailed Iguana from Utilla Island, Honduras.—No. 1217. January 19, 1901'.. 467-468 New species: Ctenosaura bakeri. On the Wheatears (Saxicola) occurring in North mMerica.—No.1220. February. 25, 1901*__.2.5-..--.-. 473-481
Watcott, Cuartes D. Cambrian Brachiopoda; Obolella, subgenus Glyptias; Bicia; Obolus, subgenus Westonia; with descriptions of new species.—No. 1229.1 May 22, Bae Oe ae wR pet eens Pe ay Po ee ae ES a ie ie Liss <5 8 669-695
New genus: Bicia.
New species: Obolella mobergi, Obolella lindstrémi, Bicia whiteavesi, Obolus (?) meneghini, Obolus tetonensis, Obolus (?) zoppi, Obolus (Lingulella) bicensis, Obolus (Lingulella) bornemanni, Obolus (Lin- gulella) linnarssoni, Obolus (Lingulella) randomensis, Obolus (Lin- gulella) schucherti, Obolus ( Lingulella) siemiradzkii, Obolus ( Acritis? ) rugatus.
New subgenera: Glyptias, Westonia.
‘Date of publication.
-LIST OF ILLUSTRATIONS.
TEXT FIGURES,
Page
Anterior portion of cranium of Trigonias osborni ........---- ed ere ate eee ee 221 SynphiysisioLjaw of Uirgonias osbornt ......2..-..2-+-22--+-+---- Eee eee Ste Dy RII ROCENIRO ONS Pent Reta ee fe one eel koe Liclcaleclocse cece tse 453 eG ACUOOMUS A maa. «sin Ste 3 on oe heys see bcecesccees SSR oy niet eet erine ech 455 OE INE EO OG Ee ae a ee a a SEE Eee Se seem seems eee ee 457 en RTICHCU SG ee Pte ec neces wh 2 Ou hee Be teal ee ee SS 461 Pere TTS GI LOCETIS IS eee a ete ty tts A eNO paren Pu ey ee Ce ot sy eer leds 464 ESOL CONIC SURE ene Sdn O Wein UE i in NG A Mahe wig oe cinwie bee aclahe 465 rani S: OROCHLCl | NOW SPCCIOS 2. 22 cfs s< 5.55 25 - sk. Me eekcc ste e ewes coecce 484 AECL se SILC Wi SICCION se ict Coe = See ee a clw sec ceteled acc eeachascsles 485 ECU ODLOWTORCl «UW, SPCCICN Sas fa c/ccicic - oa oeaa< Sele eaccalacecctceessecs 486 LLCS SOU WLC es ee es A ee 487 RetUGCICCHS IRAGIET US. NEW SPCClOS) sass s25.5h<c5eseccscl see cbse cess ceecec es 488 (COMMEP WEASEL) DO CS AG e as 5 Aa ee ee ee 489 BEN OLOCLORUS pNEW SPECClES sa. 225262 3e5 asa he. Janke de ws akes ss hece ee 491 eR PCER OMI UTOLCECWELUNO, = Seem ons Joe = as oe Se we windieieeniekenew eee 509 PRE NCUCIUL CT men Mae oe Bee he Beran en we eS 2 LE te Seles Se oe 510 PATHE TNE CCLIUT OL CLENUIONG = omens CaS 2 a Se Lees oe gece 510 Em OONOC IC UlNUTa CRERULOLGs <5 oct Reo. 2 bacco dete ele + sis encne 510 PMAGIBE TL OMMeM MONG Wir GIiand, = xat ea Ss Us. Lela ote ete ccee 512 Abdomen and last two thoracic segments of Cirolana obtruncata......---------- 514 CERCA ty enna ee en ok Ort ee ee ns = Sasol awa onda eee Saeco 515 Terminal abdominal segment of Eurydice convera........-.-------- igs eee a 516 Head and first thoracic segment of Corallana sexticornis...........------------- 518 eM MIU ON Sarma ee nae ro ose nc Jatieo a As ne laws. cee ose ee was - 526 PEG OULOMRMEAULUS, Sanaa tee cee sale cae a < os see neat Me SA ars eects een 527 Head of Meinertia transversa; abdomen of Meinertia transversa .........--------- 529 eee RECT IL vit pm ene ee Mr Pee WE ER Sova w w'c'c manne ee ence 533 Pe OMICMNOL SDNCTOMG YUCALUMMM = = oii. =- 2a eocen case ctees oe ee se esccse ences 534 Pe RE CTROLS OUNCE ONGUIIOLO —- <hr = ays s Sane eos hoe ge Senses tease ieee Ssoce 535 Head of Cilicxa carinata; lateral view of Cilicwa carinata ......-....----------- 535 ESD CLEC PN CTT OCA Soe Se 536 DEMME CLICHE TINOUNCHIAL s 326 eo nccac cout sc+de-~ cue checeee cc eoees 537 ee NOR LTIRD ATR re ro ee oe ee eee TL PS Nae 2 oe 543 SU ARURASTARCINII CITT CM TIED eer eee = eee ei ST Te) os oe ee a 544 CE el ee ene ee ere ee ee ase Uo wate be eee deans 544 PRR EE CO ciawern name sr eres oa SRL Ch seeks ee eet ale ek 547 ee NCL Pe hee see On St oe ote Se ee eee cos ous 548 PAR ELIT RECEP TUN LU UGH Sere rat et et PN Sm ay Soe Reet tyne FUE rs 502 Maxilliped of Asellus attenuatus (a); mandible of Asellus attenuatus (b); leg of
MaeMireG Pair. o.5. +. -c-< SP ee tegen ey 2 It RR TY OR Ns eS Sh 553
XII LIST OF ILLUSTRATIONS.
Page. urycope cariboed benedtel =<. s.%.. fSae sens Se oe as eee eee 559 Synuropus granulatus. Dorsal view (a); Uropoda (b); Maxilliped.........---- 563 Philoscia richmondi; Maxilliped (a); Mandible (b)...----.-------------------- 564 Sphzeroniscus portoricensis; Abdomen (a); Uropoda (inside) (b).-.--.-.-------- 573 Cancellusiornatus. 225 as5e = 2sccae cess e-ean se eeat ces ase a= See eee 772 Operculum of Cancellusionnatus .- = 32 22sec esas ee ts ee 772 Cancellus spongicola in. Sponge :< sos be os Se ee wae eee aoe oo ee ee 774 Cancellus spongicola. Extremity of abdomen ..--- --=2--2 5-255 ==Soscee- eee 774, HY/LACHICLES SPUNUUUS Soe S18 Seer aia ais ae Se al ce ee 775 Operculum ‘of Pylocheles partitus 22. 2522 sea wiee sae en ee 776 MTG DOGUIUS GUL SE Oe ee ee ec cee awe ots ean ee oe ee oe 777 Synaphobranchus Waconis 232 ae eco ernie se nee ee Oe - 845 SMO RODrANCHUS FENIING. <2 = Deine wines as Se Ri ne en ae ee 845 TAP OCEP UAVS OPED RUT <A stax m= mem re mea 850 Tee OCA UALS PUVA TUUS Se at = = Aer eee Sea tial ee ole = ae a 852 Lepiocephatus 1ystronti: 2% 52 tis ie Reese Sore Ae renee eee Sea oe ened 853 Deptocepnalus TETOURCGUS 2. = 22.2 - = ce ein See Se * 854 BOUT CLUS TLC CSU OTILUIS et a a 855 OTA UV AUS) CUCL Oe ee 856 Oxyconger leptognathts <2. 2522 fo 22320 Session cen eee ae pee ae ee 858 GChiopsis fierasper = 5% 2 ann sodas b= Lee Sie sete eee eee eee eee 860 Mureenichthys Owstont 2 2. oo. Ake ens ss scenes ee ee ee ee ee eee 862 Mairenvicnthys WOR 22 SOROS Ay pots co Names! Aeterna ss Sak ge eee 862 Marzenichthy$ Gokt so sce setae ett Jo tee ee et es Se eee 863 SPNGGeOranchus MOST’ S22 2 Pe ame tes a a alee tenet eee 222 2, OD) ‘Pisobdonophis -cophistwus. ©. 222252 23) saio ele dee eee cee eee eee ee 868 NUNIS TEUULSUS tai nc asx HR Bim Sa oe a a iaele Beet eS Re pe or I i ne 869 MiChOGOnNOD his \eTADO: Ss 22 eke <oee Bee dae se aes ee ee re ee 870 Onhichthusdasckuse (72 3 2e cee 2c nn ale cclties amor Ye gatiy bated tele Si 873 Ophichthus tauehidee 2. > 2022 522s ence 2. de seve fae ss eae be ee ee ee 873 VIN SUCAUICCNOSG: == EARS o 5 Sears ntl Si hg Se aac es as eee a ee ee ee 884 Behinda, kishtnouiet. <2 es. oso b a hese sen ose Sse see ee ee 885 Unopteryqtus ORINOWe, 02522) sa/- eas jets Bae si Sow 5-6 2s 52d ee 887 AMOGGTUCHENYS COTUNGLUS «Soci nc scn 2s alelo tees ee oe eee Skat pa ee 893 MA POGON NIGET 22 2 522 cpr k ote ee ace ie Ss et he se 895 VAOGOR, MOP GINALUS SER a = haat = Rie ae lel hm sgl aie So 897 Wpogon UNeWUges £8 J 2,5 = SS ae ee tale miata eS ere eee 899 Apogon SCRLEGEW ia/2.5 Son 2 2S okie otis eS alae oa Pac's Aral a ee i rte 900 ADOGON CCG, 222 oeinn 51a Sa he eee eine ee ele ee ee 901 AMOGON SCNIVINEOIUS = <= 52 Soa. 2 a's he cee ce ewe eae shee See eee See eee nee 903 ANOGON NOLAHUS ae os oo a ena se ee mee eee ee ae alae oe Se ee ee 904 Apogon. kiensis sa. - occos.ceccs ace ccecens-bease sees eases See ee 906 ACrODOMG JGPONICUMac oan ae eens See oe eee Be eee 912 PLATES. Facing page. 1. Venation of Tineid moths . 22.4: --2 22-22, gee ee ee ee 254 2-4. North, American Tellinida . - 20.22 sug. 36 Sma se eee ee 326 5. External aspect of right os innominatum of Basilosaurus cetoides ....---- 332 6. Internal aspect of left os innominatum of Basilosaurus cetoides .....----- 332 7. Posterior and anterior aspects of right femur of Basilosaurus cetoides ..--- 332 8. Type specimen of Leuciscus turneri ....-.----- on ace ae cee es 334 9. Gobio biwe (fig. 1); Gobio mayedex (fig. 2); Otakia rasborina (fig.3) ....-- 380 10. Ischikauia steenackeri
a
LIST OF ILLUSTRATIONS. XIII
Facing page.
EPR INILEN (I Cr Gier en es a ee Ne nti Ae ow ois = ee Se alse en see- 520 380 HOMIE UULO LUST COU GR TUBU CIE ney aaa eRe ssa sieaye aici eo a.slya as as Se's SS eee 380 SMP SAUL OLOULEREGITUIESIUUUD Were eee ene nee ale siaysiatele sioicrs ate siclae wtejsie ais.s-eeiece 380 eM SEU USLOUCSGCOCOLUS meet ear es ae eto a Son yscwwan usecase use 380 PPC MODBURCULR TOUS ee te rh malas ola Sas Seno she Sse ce Gide scree sn 380 Ree COM Cie OIG pee yates ein jaiaje aicie so ais oe ac w/a Soba ~ oss Sees s 380 SEPACILO DN TERSLOCMUCOW I, Sasa a2,- aaimatata(wciania'e se clase a Sais heecseeerecscese 380 EMTS STIS) ete eee nin 5-8 2 Semis So ciacie a ois isis ae in'as os sd eemaesas 380 RIMES LONUILUWS) 1) OLCOCUE eM eet te sisi s Ne Saniae sink Satria wime cs sine cclee o tees esse 380 UMNO OZLOTIUUIUGIUUSH GUS LUOUIY CUsetniaans Sami 8 olafe steve ale im oe she arwie: hele maw tal ay on lee ala 380 PPCM AG Ole Oleth Cm UnUTChOLNO DUS mae a= aeeis oeisen cis otc. Sareea nro (e siasterey= lore 450 Bee OMerAmiAgn anol Gerdte= ac. 5> essen aioe tae no wk cael wee e sense 590 POROUS PANE OL SICGOSHUPUS MONS. S22 o.0 «ice mece 25s =sschecemc ss 58 592 Em Oatmaliplateror Stegosaurus MATS. 5.2225 cece foto setae eee dencee ee ese 592 25-26. Skulls of Zapus, Euchoreutes, Sminthus, Dipus, and Alactaga.-..--.-.------ 668 Pye lert hind teet of Zapus, Alaciaga, and Dipus - 2-2... .--. J Ss2--422--2---- 668 PET IL CRCN SH UNCULUG HER re. 2) 2 ores re aes aaa ei win claiasisi sins Seles e Sees Scie 724 SE ORAM OMA TDRUnCUs PURCOLUS 226. Sas ak ne ek clue/Ja coe aeoeincce ot 724 RPE icte RCRSS OULU CIN oa iss as Ee asa aiala'n aeiewin co Soe See omalooe ae = Se a 734 Es OMPTAETLCTSCILSE) (O11 [ete oa ete te aye ator aes aa tao ae ote [a eral oe elena lr eee 770 Spee CL LOUIS) CD OTULGLIS ean awe oe Se ee = ele aap Sela clas ee ei sjosiawisiniss ae see 770 RMA TIC ONT MICOI OT: ame eine aera eae oleie a sons Swiacec eeeeel=eie nmin sere 770 Bee OTILUCONUTIGET. CUI LO UU Sepa ae te aye Pina stata wale ie eis nia nlaloiin ele Calle = ial 770 Sa OOH MISESITTMIS ete ciate ye omer nae acla Heb osisnwnsce ase secs Se cio an 770 DOG OOSITULUSSIUISCCULS RR tee Rye ie oa ao pata neta a (ele mafia w Siala.c afore asielae <= 770 De CLO SCORSUDICO | Cee ae eee eats ale Sate sloe eins elaine wia\Aeleie a eisai ais sie 770 SME IOTOLULGION MLO eM oe as as scare siat e cis see asses ee ones ne 5-502 qe 770 eee UN COOOL NOLLMUAMNCTICA a= - 5 oc wins toca a2 Scns sndeneccceenis~ aes chm 834
ERRATA.
Page 761. The generic name Chasmias, proposed in these Proceedings for a genus of ee gobioid fishes, is preoccupied by Chasmias Ashmead, a genus of Ichneu- ; mon insects, proposed somewhat earlier in the same Proceedings. For the genus of fishes typified by Chasmias misakius the name Chasmichthys Jordan and Snyder may be substituted.
-age 908. The generic name Melanostoma is twice preoccupied. Dr. Ginther (Deep Sea Fishes of the Challenger, XXII, p.16, 1887) has substituted for it . the name Synagrops. The Japanese species should therefore stand as Synagrops japonica. XV
CLASSIFICATION OF THE ICHNEUMON FLIES, OR THE SUPERFAMILY ICHNEUMONOIDEA.
By Witi1am H. AsHmeap,
Assistant Curator, Division of Insects.
The characters common to genera give those of the higher groups; the orders and their common characters combine to form those of the classes. It depends, therefore, wpon every classifier how far he will proceed in separation and subdivision. Indeed, much difference of opinion exists upon the determination of the groups between the species and the order, whence have arisen the several definitions of subgenus, genus, and tribe. In fact, opinions will never harmonize upon the claims of genera, because no universal prin- ciple for the structure of genera in any artificial subdivision can be given. This principle is in itself exceedingly capricious, and if one maintains thus far a genus extends, and another thus far, both are certainly right, if only every group, which they distinguish as genera, is distinguished by similar and exclusive characters. Burmeister.
The pertinency of this quotation from one of the great masters of the science of entomology will be better understood and appreciated when the body of this work is examined and it is found that no less than eleven hundred and forty genera, or more, have been recognized and tabulated, although when Burmeister penned the above lines, in 1835, the Ichneumonoidea contained only about one hundred and nine genera. It we go back to the early days of Burmeister, we find, too, that authorities differed as to the value and utility of some of these genera, just as they differ to-day. In my tables, therefore, there will be found many genera which by some eminent living hymenop- terologists are thought to be of no value, but which the writer, on the contrary, holds to be good and distinct—a difference of opinion that time alone can settle.
Very few persons have given any attention to these insects, and the necessity for these fine subgeneric and tribal divisions is evidently apparent to only a few active workers. The great majority of the workers in other groups seem totally ignorant of this vast complex, or at least have no conception of its immensity or the difficulties encoun- tered in studying and identifying material belonging to it derived from different parts of the world.
The writer has now been studying the Hymenoptera for twenty-five years, and much of this time has been devoted specially to studies in
PROCEEDINGS U. S. NATIONAL MUSEUM, VOL. XXIII—No. 1206. Proc. N. M:., vol. xxiii 1 |
2 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXIII.
the Parasitica—the Proctotrypoidea, Cynipoidea, Chalcidoidea, and, for the past ten years or more, to studies in the Ichneumonoidea. He has had material for examination from all parts of the world, and hopes, in the tables he is now publishing, to place the families, subfamilies, tribes, and genera on a better foundation, thus enabling students to avoid many of the difficulties he himself encountered, to create an interest in their collecting, and to stimulate their systematic study.
The first systematist to fully appreciate the immensity of this com- plex, to bring order out of confusion, and to lay a safe foundation for its study and classification, was Dr. Arnold Forster, of Aachen, Germany, who accomplished this great work in two contributions, entitled Synopsis der Familien und Gattungen der Braconen, pub- lished in 1862, and Synopsis der Familien und Gattungen der Ich- neumonen, published in 1868.
My own work in this superfamily is based almost entirely upon that of Férster’s, and it is scarcely necessary for me to state that with- out his contributions for my guide the present work would have been almost impossible.
The more I study Forster’s works on the parasitic Hymenoptera, the greater is my admiration for him and his work, and it was with the utmost astonishment I found that these important contributions had remained so long neglected, unappreciated, and, until within com- paratively recent years, almost totally ignored by American and European students.
Dr. Férster went too far in calling his groups families, but in the majority of cases these so-called families represented natural groups, and as such ought to have been sooner recognized. His groups in the family Braconidew have been recognized in most cases as subfamilies by the Rey. T. A. Marshall, in his monographs of the European species, while in the present work I have recognized his so-called families in the /chneumonide as either equivalent to subfamilies or tribes.
In order that the position of this immense complex in the order Hymenoptera may be thoroughly understood, I reproduce here a cor- rected table of the superfamilies:
TABLE OF SUPERFAMILIES.!
Suborder I. Heterophaga. Abdomen petiolate or subpetiolate, never broadly sessile; larvee apodous. * Hypopygium entire, and closely united with the pygidium, the sting or ovipos- itor when present always issuing from the tip of the abdomen.
'The numbering of the superfamilies and families in this paper conform to a scheme of arrangement of the whole order Hymenoptera, as proposed by the writer in John B. Smith’s Insects of New Jersey, Trenton, 1900, pp. 500-613. Tables for the recognition of the 94 families into which the order is now divided will be given at the end of this work.
No. 1206. ICHNEUMON FLIES—ASHMEAD. 2
a, Pronotum not extending back to the tegulee; trochanter one-jointed.
b. Hind tarsi dilated or thickened; pubescence of head and thorax feathery or plumose. Superfamily I. APOIDEA. bb. Hind tarsi slender, not dilated or thickened; pubescence of head
and thorax simple, not plumose. ; Superfamily Il. SPHECOIDEA.
aa. Pronotum extending back to the tegulee, or the latter absent. ce. Trochanters always one jointed.
‘d. Abdomen variable, rarely twice longer than the head and tho- rax united, most frequently much shorter; hind tibice in female neither inflated nor strongly constricted at base.
Petiole or first segment of abdomen simple, without a scale
or node; winged forms with well developed tegule. : Superfamily III. VESPOIDEA.
Petiole or first segment of abdomen with one or two scales or
nodes; winged forms without or with very imperfectly formed tegule. Superfamily IV. FORMICOIDEA.
dd. Abdomen in female greatly elongated, several times longer than the head and thorax united, the segments con- stricted at sutures and flexible; hind tibiee in female inflated and strongly constricted at base; abdomen in male not especially long, clavate. (Pelecinide. )
Superfamily V. PROCTOTRYPOIDEA (part). cc. Trochanters two-jointed.
Mandibles large, 4-dentate; hind wings with a distinct venation,
with two basal cells and a radius. (Trigonalidee. ) Superfamily III. VESPOIDEA (part). Mandibles never very large nor 4-dentate, either simple or bidentate, or at the most 3-dentate; hind wings without a dis- tinct venation, or at most and rarely with only one basal cell, the radius always absent. Superfamily V. PROCTOTRYPOIDEA. * * Hypopygium divided or never united closely with the pygidium, the oviposi- tor issuing some distance before the tip of the abdomen; trochanters always two-jointed.
d. Front wings always without a stigma, the marginal vein, if present, linear never large or stigmated; abdomen with the ventral segment hard and chitinous, without a fold.
e. Pronotum extending back to the tegule; front wings with a marginal and basal cell, either complete or incomplete; antenne straight, not elbowed.
Superfamily VI. CYNIPOIDEA.
ee. Pronotum not extending back to the tegulee; front wings
with neither a marginal cell nor a distinct basal cell,
the latter, if at all indicated, usually poorly defined by
hyaline veins visible only by transmitted light; hind
Wings without a basal cell; antennze elbowed.
Superfamily VII. CHALCIDOIDEA.
dd. Front wings with a stigma, the marginal vein usually large
or stigmated (rarely linear in some Alystidze); abdomen
with the ventral segments most frequently soft and mem-
branous, with a fold (rarely hard and chitinous without a
fold, Hvaniide and Agriotypide) ; pronotum always extend- ing back to the tegulee; antennze straight, not elbowed.
Superfamily VIII. ICHNEUMONOIDEA.
4 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXIII.
Suborder II. Phytophaga. Abdomen broadly sessile; larvee with legs; trochanters _
two-jointed. Anterior tibize with only one apical spur. ------- Superfamily IX. SIRICOIDEA. Anterior tibize with two apical spurs...-Superfamily X. TENTHREDINOIDEA.
CLASSIFICATION. Superfamily VIII. ICHNEUMONOIDEA.
This group has in the past received the following names:
1744. Ichneumon Linnzxus (part), Syst. Natur., 4th ed.
1807. Pupophaga LaTrEILLE (part), Gen. Crust. et Ins., III, p. 249.
1809. Ichneumonides, Family IV, Larrei.ie, Fam. Natur. du Régne anim., p. 444.
1823. Entomotilla, DumeErtit (part), Considér. génér. sur |. classe d. Ins., p. 220.
1837. Parasitica, eee (part), Wiegmann’s Archiy., I, p. 158.
1840. eee Diy. I, Spiculifera, WrEstwoop a Intro. Mod. Classif. Ins ul paS
1899. eae ae Superfamily VIII, AsHmMeap, Jour. N. Y. Ent. Soc., VII, p. 47.
No one, I think, who will make use of the above table of superfami- lies, can fail to place correctly any parasitic wasp belonging to this superfamily.
It is unquestionably the largest and most extensive complex in the order Hymenoptera, with possibly the exception of the Chalcidoidea, and is composed of a vast number of minor groups, representing hun- dreds of genera and many thousand species.
Not less than a million species will be found existing on the globe, although the known or described species do not yet reach much over 10,000.
Unlike some species, in others of these great complexes, all, with- out a single exception, are genuine parasites, and destroy or devour the eggs, larve, pupe, or imagoes of other insects; scarcely a single order of insects is free from their attacks, and even relatives in their own order and family are devoured by them.
The group, therefore, taken as a whole, is of the greatest economic importance, since the vast majority of the species composing it are beneficial to man. No other group of insects has a more important role in the economy of nature.
It is composed of innumerable species of the greatest variety in shape and size, from the most minute or microscopic size, measuring scarcely a millimeter in length, to forms that attain an inch, an inch and a half, or even two inches or more in length, and with or without a prominently exserted ovipositor, the ovipositor sometimes attaining a length of four or five inches, and the group is in consequence one of the most difficult and perplexing to classify.
The fauna of no single country is yet thoroughly known and our private and public collections are crowded with undescribed forms,
No. 1206. ICHNEUMON FLIES—ASHMEAD. 5
Up to the present time no general work on the group has been pub- lished, and this contribution is the first effort made to classify the group as a whole or to bring together in systematic order, or in tabu- lar form, the families, subfamilies, tribes, and the described genera of the world.
The author, although familiar with all of the groups and with most of the genera, has in some cases been compelled to rely upon descrip- tions for placing certain of the genera. The work, therefore, must be imperfect in some particulars, but he hopes for it a kindly reception, and trusts it will not only stimulate the collecting of these important insects, but that it will afford an aid and a guide for future study.
The families recognized may be distinguished by the use of the fol- lowing table:
TABLE OF FAMILIES.
Seam SI eRSBLO RIN emanate oe oes So Be eel Shs Se, SS os es ee 7 Seatac mae pee ek Sees St Ae ae i ets SS SRE Nis 3 blag 2 2. Costal and subcostal veins confluent, extending close together, side by side, threscostalecellgtheretorevabsemtween =< css eee ee oe eee eee 3 Costal and subcostal veins separated, a space between, the costal cell therefore
: present, distinct.
Abdomen inserted normally, sessile or subsessile, or the first segment long, petioliform; front wings with only one recurrent nervure; head most frequently globose and usually tuberculous. -..-......--.---- 6 Abdomen petiolated, inserted upon the dorsum of the metathorax, the body of same usually strongly compressed; front wings with one or two recurrent nervures, more rarely with none; head variable but never globose nor tuberculous; antennee 13-14 jointed, inserted either just above the clypeus or far above it on the middle of the face. Family LX XIV. Evanrp2. 3. Front wings with two recurrent neryures (the second recurrent absent only Me NESE MUSE LAr SapI0 OLESSOM) 24 22 225 2c a te'oe yest oe’ te Se 4 Front wings with only one recurrent nervure or with none -.......------- 5 4, First cubital and first discoidal cells always confluent; abdominal segments 2-3 usually flexible, rarely connate; mandibles attached normally. Ventral abdominal segments hard and chitinous, without a fold; dorsal segments 2 and 3 connate, not flexible; scutellum spined. Family LX XV. AGRIOTYPIDA. Ventral abdominal segments soft and membranous, usually with a fold; dorsal segments 2 and 3 flexible; scutellum rarely spined. Family LX XVI. IcHNEUMONID». First cubital and first discoidal cells separated, distinct, not confluent; man- dibles attached abnormally, the tips turned outwardly and not meet-
MmcuwihenGlosed== 2s a= c.o2.0-.-b see ee Family LX XVII. Atysimps. 5. Mandibles attached abnormally, the tips turned outwardly, not meeting when Close clement so eee aett rate. oe ole nee Family LX XVII. Atysip.
Mandibles attached normally.
Abdominal segments 2 and 3 most frequently rigid, connate, not flexible; if not rigid, then all the segmentsare flexible; abdomen neyer greatly elongate and strongly compressed; first cubital and first discoidal cells, although not always, most frequently distinct and separated.
Family LXNX VIII. Braconip2.
6 PROCEEDINGS OF THE NATIONAL MUSEUM. _ VOL. XXIII.
Abdominal segments 2 and 3 flexible, the abdomen very elongate, nar- |
row, and strongly compressed; first cubital and first discoidal cells always confluent. (Pharsalia Cresson.) Family LX XVI. IcHNEUMONID. 6. Antenne inserted close to the clypeus; hind femora most frequently swollen, and usually, but not always, toothed beneath. Family LX XIX. SrepHaNnip®. 7. Mandibles attached abnormally, the tips turned outwardly and not meeting whens closed... e.- Ley S=e yas) cep eee eer ere ate re ALYSUD®. Mandibles attached normally, the mandibles when closed meeting or crossing each other.
Abdominal segments 2 and 3 flexible..........-.-.----- ICHNEUMONID®. Abdominal segments 2 and 3 rigid, connate, not flexible ....BRAcoNIDz, All abdominal segments flexible (Aphidine) ......-.. BRACONID (part).
Family LXXIV. EVANIID.
1802. Evaniales LATREILLE, Hist. Nat. Crus. et Ins., III, p. 328.
1815. Evanides Leacu (part), Edinb. Eneyce., LX, p. 142.
1838. Evaniade, Family I, Hatrmay, Ent. Mag., V, p. 212.
1839. Evaniade, Family 8, Hatipay, Hym. Synop.., p. il.
1839. SHucKarp, Newman’s Entomologist, I, p. 120.
1840. Evaniide, Family 2, Westwoop, Intro. Mod. Class Ins., II, p. 124.
1883. Evaniales THomson, Opus. Ent., LX, p. 844.
1887. Evaniide Cresson, Syn. Hym. North America, p. 36.
1889. ScHLETTERER, Ann. k. k. Naturh. Hofmus., IV, p. 115.
1900. Evaniide, Family LX XIV, AsHmeAp, Smith’s Insects of New Jersey, p. 563.
This family is readily distinguished from all the others by the attach- mentof the abdomen. The abdomen is, asa rule, strongly compressed, petiolate, and attached to the dorsum of the metathorax, either just back of the scutellum or posteriorly upon or near the superior margin of the truncature, but never normally at apex, between the hind coxe, as in all other ichneumonids, with but two or three exceptions. It is further distinguished from all the other families, except the Stephanide, by having a destinct costal cell in the front wings, the costal and sub- costal veins, unlike other ichneumonids, being distinctly separated.
The family is usually divided into two subfamilies, but I have here recognized three major groups, separable upon good structural char- acters, and further supported by their economy or different habits of the species composing them.
These three subfamilies may be easily recognized by the aid of the following table:
TABLE OF SUBFAMILIES.
Antenne inserted far anteriorly just above the clypeus...............-i2------- 2 Antenne inserted far above the elypeus on, or very near the middle of the face. Front wings without or at most with only one recurrent nervure; venation in hind wings wanting or indistinct, without a median cell. Pronotum very short, transverse linear and abruptly truncate anteriorly; abdomen attached by a petiole to the superior margin of the metatho-
Clin whoo tng a > Sl die hi ae
NO. 1206. ICHNEUMON FLIES—ASHMEAD., ,
racic truncature, remote from the scutellum, the body short and com- pressed, fhe ovipositor not or at most subexserted; head viewed from
above transverse, the temples never very broad. Subfamily I. evanun x. Pronotum elongate, conical, never transverse linear, abdomen attached to the base of the metanotum just behind the scutellum, the body very long, usually long, sickle-shaped, compressed; the ovipositor long or always strongly exserted; head viewed from above subtriangular or obtrapezoidal, the temples oblique but very broad or broad, more rarely ploboseieners jason te ee set cats Subfamily I]. GAsTERUPTIONIN®. 2. Front wings most frequently with two recurrent nervures, the second sometimes subobsolete, rarely wholly absent; hind wings with a distinct median cell; abdomen clavate, not strongly compressed, the ovipositor exserted. Subfamily III. auvnacinz.
Subfamily Il. HVANIIN 2. 1900. Evaniine, Subfamily II, Asameap, Smith’s Insects of New Jersey, p. 563.
The position of the antenne, the venation of the front and hind wings, as well as the shape of the abdomen, readily distinguish this group from the Au/acinw, while from the Gasteruptionine it is at once separated by the shape of the head, the very short truncate pro- notum, and the short, strongly compressed, hatchet-shaped abdomen and its attachment to the metathorax.
All the species falling in the subfamily are parasitic in the eggs of cockroaches. Lvania appendigaster Linneus, a species now widely distributed to all parts of the world, has been frequently bred from the eggs of these insects. In Florida I have reared it from the eggs of Periplaneta americana Linneus and P. australasie Fabricius. I have also a specimen of //yptia dorsalis Westwood, bred by Mr. Weed, in Mississippi, from the eges of /schnoptera pennsylvanica De Geer.
Only two genera are known, distinguished as follows:
TABLE OF GENERA. Front wings without a marginal cell and also without cubital and discoidal cells. (1) Hyptia Illiger. Front wings with amarginal cell and also with one or two discoidal cells. (2) Evania Fabricius.
Subfamily II. GASTERUPTIONIN 4.
This group, or subfamily, is at once separated from the Avlacine by the insertion of the antenne, the venation of front wings, and by the attachment of the abdomen, which is joined to the metathorax just behind the scutellum.
It approaches nearest to the Lvan7zinw, but is easily separated by the quite different shaped head, which is long, obtrapezoidal, as viewed from above; by the very long conical pronotum; by the abdomen, which is very long, narrow, and strongly compressed, and attached
8 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXIII.
differently, and by the shape of the hind legs, which differ decidedly from the other two subfamilies, the femora being shorter and thicker, the tibie very strongly clavate, while the basal joint of the tarsi is stout, and as long or a little longer than the following joints united: The habits of the species, too, are quite different from the others, since all whose parasitism is known have been bred from the nests of wasps and bees—Cvrabro, Philanthus, Cerceris, Gorytes, Stizus,
Eumenes, Odynerus, Sphecodes, Prosopis, Halictus, Andrena, ete.
The two genera falling in this group may be separated as follows: TABLE OF GENERA.
Front wings without a distinct venation, at most with only slight traces of a venation, as in Pelecinus; head globose and deeply excavated anteriorly above for the reception: of the scape i752-2<¢ 2255 see ee re ee (3) Leptofoenus Smith.
Front wings with a distinct venation; head large, viewed from above subtriangular or obtrapezoidal, the temples oblique, broad; no excavation anteriorly for the reception of the scape......----- (4) Gasteruption Latreille = Foenus Fabricius.
Subfamily III. AULACIN 2“.
1840-42. Aulacidx, Family, SHuckarp (part), Newman’s Entomologist, p. 121. 1900. Aulacine, Subfamily I, Asomeap, Smith’s Insects of New Jersey, p. 563.
This group was first recognized by W. E. Shuckard as above, but he incorrectly included as components of it Zr/gonalys Westwood and Lycogaster Shuckard, which have no real affinity with it, but represent a distinct family far removed from any family belonging in this series.
The Zrigonalide are now placed in the superfamily Vespotdea between the Lethylide and the Sapygide.
The Awlacine, as here restricted, are easily distinguished from the other two subfamilies by having the antenne inserted on the anterior margin of the head, just above the clypeus, by the quadrate or sub- globose head, and by the venation of the front wings, which have usually two recurrent nervures.
The abdomen, too, is quite different from the other groups, being elongate, clavate, and only slightly compressed.
All of the species are parasitic on the larvee of different Coleoptera, those belonging to the family Cerambycide being particularly subject to their attacks.
Three genera have been recognized, distinguishable as follows:
TABLE OF GENERA.
First cubital cell receiving the first recurrent neryure toward the middle; hind coxee swollen, much elongate, and prolonged within beyond the insertion of thestrochanterss= 228s Joo es Sea eee (5) Pammegischia Provancher.
First cubital cell receiving the first recurrent nervure at or near the tip, or inter- stitial with the first transverse cubitus; hind coxe normal, not prolonged
within. Clawsiwithione toothy wemeat ieee ees ee ee (6) Aulacus Jurine. Claws with three or more teeth beneath................ (7) Pristaulacus Wieffer.
No. 1206. ICHNEUMON FLIES—ASHMEAD. 9g
Family LXXV. AGRIOTYPID.
1832. Agriotypus WALKER, Curtis Brit. Ent., IX, pl. 389.
1888. Agriotypide, Family III, Haurpay, Ent. Mag.,V, p. 212.
1868. Agriotypoide Forster, Verh. d. naturh. Ver. pr. Rheinl., X XV, p. 143. 1884. Agriotypide BripGMaAN and Fircn, The Entom., X VII, p. 121.
This is probably one of the most interesting families in the super- family /chneumonoidea, not only on account of its rarity, its structural peculiarities, and its aquatic habits, but also on account of being rep- resented, up to the present time, by but a single genus with a single species—the Agriotypus armatus Walker.
A doubt as to its proper position in this great complex has been expressed by several eminent entomologists, and quite recently Dr. David Sharp has suggested its close relationship with the Procto- trypide. I myself have long had doubts as to its true position, but now, after a careful study of specimens of both sexes, am prepared to defend its position here, the only character at variance with any in this major group being in the abdomen, which has the venter hard and chitinous, as in the higher Hymenoptera, the aceu/eata or monotrocha. All its other characters are, however, as with the genuine Ichneumo- nids and with the Braconids.
The trochanters are two jointed; the wingsand their venation as in most Ichneumonids, the costal cell being absent, the subcostal vein lying close to and extending parallel with the costal vein; there are two recurrent nervures, the second received beyond the first transverse cubitus, the first cubital and first discoidal cells confluent, the first abscissa of the cubitus being absent; there are two basal cells, two complete discoidal cells, and a short triangular marginal cell, while the stigma is broad and oblong; the hind wings have a distinct venation, the submedian cell being about half the length of the median, the subdiscoidal nervure being distinct and originating from the transverse median nervure a little below the middle.
The venter, although hard and chitinousas in the genuine wasps and Proctotrypids, has the ovipositor subexserted, issuing from before its tip, and structurally is the sameas in the Ichneumonidsand the Braconids. In the male the external claspers are unusually long and broad, a char- acter sometimes met with in males belonging to the 7ryphoninw and the Ophionine. The spined scutellum in Agriotypus is quite unique, al- thougha somewhat similar spined scutellum is found in some Ophionines.
The only species, Agriotypus armatus Walker, is unknown outside of the European fauna. It attacks the larve of various species of Trichoptera belonging to the genera Silo, Goera, Trichostoma, Aspa- therium, and Odontocerum and has been observed swimming and diving under water to seek its prey.
Generic-characters same as family -.-..(1) Agriotypus Walker = Cratopus Holmgren. (Type, Agriotypus armatus Walker. )
10 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXUI,
Family LXXVI. ICHNEUMONIDZ.
1815. Ichneumonida Leacn (part), Edinb. Encycl., TX, p. 142.
1837. Parasitica Hartia (part), Wiegmann’s Archiv., I, p. 158.
1838. Ichnewmonidx, Family II, Hatipay, Entom. Mag., V, p. 4.
1840. Ichneumonidx, Family III, Wesrwoop, Intro. Mod. Class., Ins., IT, p. 83. 1900. Ichneumonide, Family LX XVI, ASHMEAD, Smith’s Insects of New Jersey.
This family is readily distinguished from the Avaniide and the Ste- phanide by the absence of a distinct costal cell in the front wings, the costal and subcostal veins being parallel and extending close together, side by side, to the stigma; by the abdomen being attached normally, not high up on the dorsum of the metathorax, and by the venation of the hind wings. From the Alysz7d@ it is separated by the normally attached mandibles, as well as by palpial characters, while from the Braconidae it is separated by the venation of the front wings, having, except in a single case, two recurrent nervures, whereas the Braconidw have none or only one. The first cubital and the first discoidal cells are also always confluent, not distinctly separated as in the normal wings of a Braconid, and also by the usually longer abdo- men and by the flexibility of the first and second segments, which in the Braconidae are rigid, connate, or not at all flexible, except in the subfamily Aphidiine.
The family /chneumonide may be divided first into five major groups, called subfamilies, as follows:
TABLE OF SUBFAMILIES.
First abdominal segment straight, not elbowed, most frequently sessile or subsessile, more rarely petiolate, its spiracles usually placed at or before the middle, more rarely somewhat behind the middle; in the latter case the abdomen is compressed; if petiolate, the petiole is usually abruptly enlarged at apex, the spiracles being closer to each other than to the apical margin (very-farely widely separated) 2. ..2..s2 32 2eseasdee cess] 2
First abdominal segment petiolate, not straight, or very rarely, but depressed, curved, bent, or elbowed, and most frequently widened at the apical third, its spiracles placed always beyond the middle or between the middle and the apex; areolet in front wings usually pentagonal or small quadrate, rarely deltoid, petiolate, or rhomboidal, although often absent.
Mesosternum not separated from the mesopleura by a grooved line or furrow; spiracles of first abdominal segment wider from each other than to the apex of the segment; ovipositor hidden or at most subexserted; areolet of front wings pentagonal, rarely deltoidal or rhomboidal, or incomplete; TO PLCTOUSHOnMIS(s =. a= ee Subfamily I. 1coNEUMONIN®.
Mesosternum separated from the mesopleura by a grooved line or furrow; spiracles of first abdominal segment nearer to each other than to the apex of the segment; ovipositor exserted, prominent, rarely very short; areolet of front wings pentagonal or small quadrate, often incomplete or wanting; apterous and subapterous forms common.
Subfamily II. cryprinz.
2. Abdomen usually depressed and sessile, never strongly compressed, although
sometimes compressed toward apex, more rarely petiolate; spiracles of
a
ee eS ee
NO. 1206. ICHNEUMON FLIES—ASHMEAD. be
first segment placed at or a little before the middle, -rarely slightly behind the middle.
Abdomen elongate, subcylindrical, most frequently sessile, rarely petiolate or subcompressed at apex; ovipositor always prominent, often very long; areolet in front wings, when present, usually rhomboidal or tri- angular, very rarely pentagonal ......-.---- Subfamily IIT. prrerin x.
Abdomen not or rarely very long, depressed, and sessile, fusiform, clavate, ovate,-or oval, more rarely distinctly petiolate; ovipositor hidden, never prominent, at the most subexserted; areolet triangular, rhom- boidal or wanting, rarely pentagonal - .-.-- Subfamily IV. rTrRyPHONIN®.
Abdomen usually long, wholly compressed or compressed along the posterior half, rarely subcylindrical; in the latter case the petiole is somewhat abruptly dilated at apex; spiracles of first segment most frequently placed at or behind the middle, more rarely before; areolet in front wings usually triangular, rhomboidal or wanting, often petiolate; ovi- positor either hidden or prominent ----.---- Subfamily V. opHIONIN».
Subfamily I. ICHNHUMONIN . 1900. Ichneumonine, Subfamily I, AsHmEap, Smith’s Insects New Jersey, p. 563.
To this subfamily belong Forster’s families Zrogotde (= Joppine Kriechbaumer), /chneumonoide, Listrodromoide, Alomyoide, and Pheogenoide, which, however, are here recognized as tribes, since they represent natural minor groups.
The tribes recognized in this subfamily may be separated by the use of the following table:
TABLE OF TRIBES.
Metathoracie spiracles round or circular, more rarely broadly oval; claws simple, TLE Lae OC CUA ALE Wappen erie eee ae ee aS Ao chee Bape aS eyecare Siar 3 Metathoracic spiracles linear or long-oval, but very rarely rounded; if rounded the claws are always pectinate. Mandibles not bidentate, simple, edentate, acute at apex....---------------- 2 Mandibles bidentate; head not broader than long.
Metanotum with a strong constriction or furrow between it and the post- scutellum, the metanotum usually short, with a median elevation toward base and without the basal or first median area, or, if at all present, open; areola often reduced to a tubercle, or if defined usually confluent with the petiolar area, rarely distinct, horse-hoof shaped or broadly transverse; scutellum variable, frequently cone- shaped, pyramidal, or highly convex, rarely very flat; sutures between the abdominal segments often strongly constricted; areolet in front wings tetragonal, triangular, or pentagonal (rarely wanting).
Tribe I. Joprrnt.
Metanotum without such a constriction or furrow, at most with only a weak furrow between it and the postscutellum; metanotum rarely short, always without a median elevation at base and with the basal or first median area distinct, usually complete, the areola and petiolar are separated, distinct, abdomen normal, not or rarely strongly con- stricted between the segments; areolet in front wings pentagonal.
Claws simple; second and third abdominal segments with lunulee. Tribe II. IcHNEUMONINI. Claws pectinate; second and third abdominal segments most frequently without lanulee or at least not present on both segments. Tribe III. Lisrropromint.
12 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL, XXIII.
bo
Head, viewed from in front, broader than long; occiput strongly concave, the temples: broad, full. 3... 23-2 22-ssesees Tribe IV. HEeREsIARCHINI. 3. Metanotum without the basal or first median area, the areola fully two and a half times as long as wide and acutely pointed at base; petiolar area not longer than wide; metathoracic spiracles large, broadly oval. Tribe V. ALomyINtI. Metanotum with the basal or first median area distinct, the areola never twice as long as wide and not pointed at base, either truncate or rounded, or, at the most, obtusely triangular at base; petiolar area much longer than wide; metathoracic spiracles rounded or circular, never Taree sei vec cc selec tle eee Ree ee ae eee Tribe VI. PH2x0GENINI.
ripe. Orme Tie
1868. Trogoide, Family 27, Férsrer, Verh. d. naturh. Ver. pr. Rheinl., XXV, pp. 144 and 188.
1894. Trogini, Tribe I, AsHmpap, Proc. Ent. Soc. Wash., III, p. 278.
1898. Joppine, Subfamily, Krrecupaumer, Ent. Nachr., XXIV, p. 2.
1900. Joppini, Tribe I, AsHMEAD, Smith’s Insects of New Jersey, p. 563.
The typical forms falling in this tribe are easily separated by anten- nal, scutellar, and abdominal characteristics from those falling in the tribe Jchneumonini; but there are several genera which can scarcely be distinguished from genuine ichneumonini, and these must be exam- ined carefully for the metathoracic characters made use of in my table of tribes.
Forster based the group upon the genus Zrogus Gravenhorst and gave for the family diagnosis a single character—the shape of the scutellum. Kriechbaumer has done no better, although he calls the group a subfamily—the Joppine, taking the name from the first- described genus, Joppa Fabricius. He has, however, given an ex- cellent table, and brought together a number of genera closely related. Kriechbaumer does not include in his table /oppa Fabricius, but for the species usually considered as such he has proposed the name Microjoppa. He had, of course, the right to segregate the old genus Joppa, but no right to reject it entirely and I have here restored the name Joppa for his Microjoppa. His genus Tricyphus, too, seems to be identical with 7rogus Gravenhorst.
The following table will aid in separating the genera belonging to this tribe:
TABLE OF GENERA.
Antennie in female at or beyond the middle widened, compressed; in male beneath serrate; abdomen with more or less distinct angular emargina-
tions, the segments, at least in part, striate or aciculate....-- 2 Only one or the other characteristies pres@ht._. 522-22 2220-2 22 eee ee eee 6
2. Wings with at least the tips brown, often also with brown maculee or bands toward the base or before the middle, seldom wholly or in great part browil. 2 Suess2s0c66 Soe eee eee
Wings hyaline, the widening of the antennz often insignificant, scarcely perceptible: 2 :2s4.og8 22 oe es ee eee ee eee eee 4
3. Labrum hidden under the clypeus; head large, with the cheeks more or less
swollen; species rather small, mostly yellow, or reddish yel-
ah eal od
| | |
No. 1206. ICHNEUMON FLIES—ASHMEAD. 13
low and black; areolet in front wings usually oblique, trape- zoidal, not petiolate; scutellum convexly rounded; gastrocceli distimets. = 2. : (1) Joppa Fabricius= Microjoppa Kreichbaumer. (Type, Joppa dorsata Fabricius.) Labrum prominent or projecting; species rather large.
Fifth dorsal abdominal segment in female inclosing the sixth; in male the sixth inclosing the seventh; apex in both sexes sometimes extending into a short point----(2) Cryplopyge Kriechbaumer.
(Type, Joppa picta Guérin. )
Fifth dorsal abdominal segment in the female and the seventh in the
male distinctly visible; areolet oblique, trapezoidal, petiolate.
(3) Macrojoppa Kriechbaumer.
(Type, Joppa blandita Cresson. )
4. Antenne in male lengthened, but scarcely perceptibly widened; areolet pentag-
O11 ee eters ee alee are eerie ease ake Eee 5
Antenne in male much thickened at the middle or strongly widened and again
narrowed; abdominal segments two and three very flat and
with fine close aciculations, the following compressed, almost
conically pointed, with fine scattered punctures; areolet small,
pentagonal, but so narrowed and contracted as to appear nearly
Tetrac OMe pe see ea eee en Sea (4) Conopyge Kriechbaumer.
(Type, Conopyge cinctipes Kriechbaumer. )
5. Scrobes normal, the lateral margins not produced into tubercles; gastrocceli distinct.
Basal joint of hind tarsi produced below into a flattened leaf-like projection.
(5) Ileanta Cameron.
(Type, Zleanta latitarsis Cameron. ) Basil joint of hind tarsi normal.
Labrum prominent, projecting; aciculations of abdomen in male very strong, extending to the middle of the fourth segment, in female Onetokune diithe = seen ease (6) Lindigia Kriechbaumer. (Type, Lindigia varia Kriechbaumer. ) Labrum hidden under the clypeus. Abdomen with the aciculations extending only to the middle of thethird sepment-&. 62.4.2 2: (7) Pecilojoppa Kriechbaumer. (Type, Peecilojoppa histrio Kriechbaumer. ) Abdomen with all the segments aciculated; scutellum margined at sides; submedian cell a little shorter than the median. (8) Ortezia Cresson. (Type, Joppa egregia Cresson. ) Abdomen with segments two and three ruguloso—punctate, the following almost smooth, shining; gastrocoeli large, oblique, deep, with a narrow space between; scutellum convex; abruptly declivous posteriorly, the sides margined; metathorax with the upper hind angles briefly dentate, the areolet present. (9) Henicophatnus Kriechbaumer. (Type, Henicophatnus rufithorax Kriechbaumer. ) Scrobes with the lateral margins produced into slightly curved tubercles; gastrocceli wanting; scutellum flat; abdomen strongly punctate. (10) Abzaria Cameron. (Type, Abzaria latipetiolaris Cameron. ) 6. Antenne in female dilated between the middle and the apex, more rarely scarcely perceptibly dilated; abdominal segments truncate, without dis- tinct aciculations or foveate impressions, usually punctate; male often difficult to separate from those in the Ichneumonini; areo-
14 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXIII.
let trapezoidal, rarely quadrate, triangular, or pentagonal (rarely wanting))\ 22o2/s2.20 <3 ce oa aalte ee ee ee oe ao Antenne in both sexes filiform, not perceptibly dilated at the middle. Abdomen with more than three visible dorsal segments, and aciculate or striate: s..o Sess cn Fae Oe a ee eee a Apdomen with only three visible dorsal segments, closely and strongly punc- tate, the third at apex ending in a strong tooth on each side; scutellum at apex tridentate ....-...- (11) Rothneyia Cameron. (Type, Rothneyia wroughtonii Cameron. ) 7. Legs long and slender, the hind femora extending to or beyond the tip of the abdomen; last ventral segment entire; shape of body and color of wings similar to Macrojoppa species. (12) Ischnopus Kreichbaumer. (Type, Ischnopus longiceps Kreichbaumer. ) Legs shorter, at the most the hind femora extending only to the apex of the fourth dorsal segment. Scutellum flat; wings wholly violaceous black or the anterior are marked witht yelllow=2See.o-eeeee oee (13) Pedinopelte Kreichbaumer. (Type, Joppa Gravenhorstii Guérin. ) Seutellum subquadrate, truncate at apex, subconvex above and margined laterally; metathorax short, imperfectly areolated, the spiracles elongate linear; areolet in front wings triangular. (14) Obba Tosquinet. (Type, Obba celatus Tosquinet. ) | Scutellum more or less pyramidal or conical, immargined; areolet in front wings subpentagonal or subrhomboidal. (15) Dinotomus Forster = Psilomastix Tischbein. (Type, Ichneumon lapidator Fabricius. ) 8. Seutellum elevated, convex, conical or saddle-shaped; posterior face of meta- thorax with three parallel areas, rarely entirely wanting or undistinetly defined... 'U. seit ees as ee 9 9. Metathorax normal, the upper hind angles not produced into teeth or spines. .10 Metathorax with the upper hind angles produced into teeth or spines, or with a very shanpiledgve ts 222,22. Se. fo ea eee 14 10. Areolet small or only moderately large, trapezoidal, triangular, or pentagonal. -.11 Areolet large, in outline quadrate; abdomen with normal number of segments. (16) Tetragonochara Kriechbaumer. (Type, Joppa polychroa Brulle. ) Areolet wanting; abdomen with only three visible dorsal segments. (17) Chreusa Cameron. (Type, Chreusa fulvipes Cameron. ) Jae Abdomen wii/ianventralyslituiapexses-ssee soe sass ae ae eee 12 Abdomen without a ventral slit at apex. 5 ’ Scutellum and metanotum at base elevated, the postscutellum between also sometimes with a small elevation, the metanotum very short, obliquely truncate from near base; areolet in front wings trape- zoidal or rhomboidal, more rarely pentagonal.
Labrum hidden, areola of metanotum obsolete or very minute, some- times represented by a tubercle; areolet in front wings not pen- tagonal; scutellum subconical, not margined laterally.
(18) Trogus Gravenhorst = Tricyphus Kriechbaumer.
Labrum not entirely hidden, rounded anteriorly; areola of metanotum distinct, usually horse-hoof shaped; areolet in front wings usually pentagon a) es ae ee (19) Automalus Wesmael.
(Type, Trogus alboguttatus Gravenhorst. )
‘ No. 1206. ICHNEUMON FLIES—ASHMEAD. v5
Scutellum flat or subeonyex, the metanotum not elevate, the areola distinct; female antennee slightly flattened beyond the middle; metano- tum not short. |
Areola horse-hoof shaped, a little longer than wide; basal lateral and middle lateral areas confluent ---(20) Protichnewmon Thomson. (Type, Ichneumon fusorius Linnzeus. ) Areola not distinctly horse-hoof shaped, a little wider than long; basal lateral and middle lateral areas separated. 3 (21) Ceelichneumon Thomson. Type, Ichneumon lineator Grayenhorst. fieaAntenne in female yery slightly widened -............-........-.----.--- 13 Antennze in female distinctly lanceolate.
Areolet tetragonal, pyramidal; metathorax very short, strongly declivous;
thesaneolaiverysshort 5. =. 22.2522. - (22) Catadelphus Wesmael. (Type, Ichnewmon arrogator Fabricius. )
Areolet irregularly pentagonal or nearly trapezoidal, the veins sometimes curved, as in Dinotomus Forster.
(23) Camarota Kriechbaumer. (Type, Camarota thoracica Kriechbaumer. )
Areolet pentagonal; scutellum margined laterally and posteriorly; head
almost quadrate; abdomen narrow. (24) Ischnojoppa Kriechbaumer. (Type, Joppa lutea Fabricius. ) 13. Scutellum convex, with lateral ridges at base; areolet irregularly pentagonal; metathorax areolated; abdomen strongly punctate, the seg- ments 2-5 constricted at the sutures. (25) Trogomorpha Ashmead, new genus. (Type, Ichneumon trogiformis Cresson. ) Scutellum saddle-shaped; i. e., pyramidal, with an emargination at apex; meta- thorax exareolated; abdomen aciculate and rugulose. (26) Microsarge Kriechbaumer. (Type, Microsarge sieberi Kriechbaumer. ) Scutellum cushion-shaped, surrounded by a distinct, elevated margin, the field thus formed nearly horse-hoof shaped; metathorax exareolated, the hind angles rounded, with only a small tubercle; abdomen finely punctate-rugulose; all tarsi long, as long or a little longer than their femora. Female...(27) Hoplojoppa Kriechbaumer. (Type, Hoplojoppa parvispina Kriechbaumer. ) Seutellum thorn-shaped; metathorax areolated as in Ichneumon; abdomen rather flat, subclavate, finely and moderately regularly acicu- late and rugulose; female antennz scarcely perceptibly wid- CM CGaNie Aefs hues Lee ee (28) Stenolonche Kriechbaumer. (Type, Stenolonche areolata Kriechbaumer. ) 14, Labrum prominent, distinct; metathoracic spines very large; scutellum flat and margined to beyond the middle, the margins anteriorly acutely elevated; areolet pentagonal; submedian cell a little longer than the median, the disco-cubital vein broken at the middle by a slight stump of a vein..-.. (29) Cryplojoppa Kriechbaumer. (Type, Cryplojoppa semicastanea Kriechbaumer. ) Labrum hidden; metathoracic spines small.
Head transverse, the temples not especially broad; scutellum saddle-shaped, emarginate above; areolet pentagonal or nearly trigonal; anten- nee feebly dilated...-..------ (30) Eccoptosarge Kriechbaumer.
(Type, Eccoptosarge Waagenii Kriechbaumer. )
16 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXIII.
Head large, swollen, the occiput deeply concave; scutellum very broad, subquadrate, more or less elevated, and margined at the sides, unituberculate, or with a small spine above; areolet rather small, pentagonal, the median and submedian cells of an equal length; gastrocceli very large transverse.
(81) Gdicephalus Cresson. (Type, Gdicephalus longicornis Cresson. )
Head transverse, or subquadrate; scutellum cushion-shaped, convex, and
margined; areolet trapezoidal; metathorax exareolated. (Male). (27) Hoplojoppa Kriechbaumer. (Type, Hoplojoppa parvispina Kriechbaumer. )
Tribe ll. [CHNEUMONINE
1868. Ichneumonoidx, Family 29, Forstper, Verh. d. naturh. Ver. pr. Rheinl., XXYV, pp. 144 and 149.
1894. Ichneumonini, Tribe II, AsHmMgEap, Proc. Ent. Soc., Wash., III, p. 278.
1900. Ichneumonini, Tribe II, ASHMEAD, Smith’s Insects of New Jersey, p. 564.
As previously stated, this tribe is scarcely separable from some forms belonging to the Joppinz, and it requires considerable care and the closest scrutiny for the detection of the metathoracic differences, used in my table of tribes, before one can be sure of the position of certain forms. It is clearly connected with the Joppini by the genus Amblyteles and allied genera through Protichneumon, Calichneumon, and Automalus.
The tribe is, however, easily separated from the others: The simple, non-pectinate claws separate it from the Listrodromini, the bidentate mandibles from the Heresiarchini, while the large, elongate or linear spiracles distinguish it from the Alomyini and the Phwogenini.
The genera may be distinguished by the use of the following table:
TABLE OF GENERA.
Basal third of petiole flattened, wider than thick dorso-ventrally ........-----.-- 8
Basal third of petiole not flattened, or so little that it is not wider than thick dorso- ventrally.
Abdomen in female with its tip blunt, the last ventral segment covering the base
of the: borers (Ambiypy qt) oo aeons ee eee 5 ~ Abdomen in female with its tip pointed, not blunt, the last ventral segment not covering the base of the borer (Oxypygi)-.----------------- 2
2. Scutellum not short, convexly elevated and declivous posteriorly, not margined laterally; metathorax with the upper hind angles usually den- tate, the areola wider than long.
(32) Hoplismenus Grayenhorst. (Type, Hoplismenus perniciosus Gravenhorst. ) Scutellum short, subconvex, rounded posteriorly and margined laterally, meta- thorax unarmed, the areola narrow, curved and much broader than long 2.35 beSseq gene eee eee (83) Callimus Tosquinet. (Type, Callimus adornatus Tosquinet. )
Scutellum usually flat, never much elevated nor highly declivous posteriorly. Clypeus medially on the anterior margin not emarginate or sinuate, either truncate or slightly rounded *2222-=eeee a eee 3
ih ated
ee ney ee ee aR See ee
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i act ci Sl
cfd.
No. 1206. ICHNEUMON FLIES—ASHMEAD. i
Clypeus medially on the anterior margin, emarginate or sinuate; metathorax with the areola elongate rectangular, the labrum more or less exposed; ciliate; antenn:e filiform.
(84) Chasmias Ashmead, new name. = Chasmodes Wesmael nec Cuvier et Valenciennes. (Type, Ichneumon notatorius Grayenhorst. ) 3. Ovipositor and sheaths not or only slightly extending beyond the tip of the PUM OMEN eee Renae ee rae et cacti ce cee een ea eens 4 Ovipositor and sheaths thickened and extending beyond the tip of the abdomen. Antenne filiform; metathorax with the areola large, nearly hexagonal;
: eighth dorsal abdominal segment exserted. (35) Exephanes Wesmael. (Type, Ichneumon hilaris Grayenhorst. ) 4. Second abdominal segment cask-shaped, the sutures between segments 2, 3, andlAsvenysdeep= os. eo a. soe aa (36) Pithotomus Kriechbaumer. (Type, Pithotomus rifiventris Kriechbaumer. ) Second abdominal segment normal, not cask-shaped, trapezoidal, or rectangular.
Abdomen subdepressed, the petiole feebly bent.
’ (37) Diphyus Kriechbaumer = Diphyes Kriechbaumer. (Type, Diphyes tricolor Kriechbaumer. ) Abdomen convex, the petiole strongly curved or bent at the posterior third; Anterior tarsi in female somewhat dilated-.-(388) Hupalamus Wesmael. (Type, Eupalamus oscillator Wesmael. )
Anterior tarsi in female normal.
Areola of metanotum quadrate or nearly, the basal lateral and the middle lateral areas confluent; post petiole scabrous or rugulose; flagellar joints 2-4 in female three or more times longer than thick.
(39) Stenichnewmon Thomson. (Type, Ichneumon pisorius Linnzeus. )
Areola of metanotum quadrate, usually a little longer than wide, the hind margin curved inwardly or more or less angularly emarginate, the basal lateral and the middle lateral areas usually, but not always, separated; post petiole aciculate; flagellar joints 2-4 in female short, scarcely or not much longer than thick.
(40) Ichneumon Linneeus. (Type, Ichneumon luctatorius Linnzeus. )
Areola of metanotum large, hexagonal or subquadrate, the basal lateral and the middle lateral areas usually separated; post petiole punctate; flagellar joints 2-4 in female subequal, longer alriat lat keseeett: Sharer Ne eo (41) Melanichneumon Thomson.
(Type, Ichneumon spectabilis Holingren. )
Areola of metanotum horse-hoof shaped or nearly, a little wider than long, or cordate, the basal lateral and the middle lateral areas complete; antennze in female stout; flagellar joints 2-4 longerithanithi ck saeco = =] (42) Cratichneumon Thomson.
(Type, Ichnewmon luteiventris Thomson. )
Areola of metanotum nearly semicircular, wider than long, the basal lateral and middle lateral areas separated; head subquad- rate; antenne and legs stout; flagellar joints 2-4 in female quad- rate or nearly, not or scarcely longer than wide.
(43) Barichneumon Thomson. (Type, Ichneumon anator Gravenhorst.)
2
Proc Ns M.> vol: xxii
18 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXIII. 5. Abdomen in female very long and much compressed toward apex. (44) Limerodes Wesmael. (Type, Limerodes ophionoventris Wesmael. ) Abdomen in female neither especially long nor compressed toward apex. Abdomen in female with 7 dorsal segments; joints 12-16 of male antennze mot! widened = eoe2 2: = =. See eee eo eee 6 Abdomen in female with 8 dorsal segments; joints 12-16 of male antennze somewhat widened. Scutellum normal. Metathorax unarmed, the spirales oval; abdomen very slender. (45) Hypomecus Wesmael. (Type, Hypomecus albitarsis Wesmael. )
Metathorax normal, bispinose, or bidentate, the spirales elongate or linear; abdomen not slender; male antennz slender, the joints nodulose beneath.
Abdomen without ventral fold, except sometimes on first segment; gastroceeli and thyridia large, deep, broader than the space between; seventh segment in both sexes black; areola of metanotum in outline circular with its apex truncate.
(46) Clenichneumon Thomson. (Type, Amblyteles funereus Gravenhorst. )
Abdomen with ventral fold on segments | and 2 or1 to 3; gastrocceli and thyridia small or moderate.
Mesosternal epicnemia entire; dorsal abdominal segments 6 and 7 spotted with white or yellow; hypopygium large, almost entirely covering the terebra; third ventral segment rarelyvawitoa olds ee (47) Spilichneumon Thomson.
(Type, Amblyteles occisorius Gravenhorst. )
Mesosternal epicnemia not entire; anus usually pale; hypopy- gium usually not attaining the terebra; third ventral seg- ment usually with a fold.
Upper hind angles of metathorax unarmed. (48) Pseudamblyteles Ashmead, new genus. (Type, Amblyteles paliiatorius Gravenhorst.) Upper hind angles of metathorax distinctly bispinose or bidentatessase see es eee (49) Amblyteles Wesmael. (Type, Ichneumon bidentorius Fabricius = fasciatorius Wesmael. ) Scutellum gibbous; metathorax bidentate; abdomen short, oval. (50) Hybophorus Kriechbaumer. (Type, Ichneuwmon aulacus Gravenhorst. ) 6. Pronotal furrow normal, not interrupted medially by an elevation or keel... 7 Pronotal furrow interrupted medially by an elevation or keel. (51) Anisobus Wesmael. (Type, Ichnewmon cingulatorius Wesmael. ) 7. Tarsi on the underside pilose, without or with very small spines. Metathorax with the areola longer than wide. (52) Hepiopelmus Wesmael. Type, Ichneumon leucostigmus Gravenhorst. ) Tarsi on the underside pilose, with strong spines. Clypeus anteriorly strongly rounded and medially toothed or angulated. (53) Acolobus, Wesmael.
(Type, Acolobus sericeus Wesmael.) —
No. 1206. ICHNEUMON FLIES—ASHMEAD. 19
Clypeus anteriorly straight, truncate. Scutellum quadrate; antennal joints 12-16 dilated laterally. Male. (45) Hypomecus Wesmael. (Type, Hypomecus albitarsis Wesmael. ) Scutellum not quadrate; antennal joints 12-16 not dilated laterally.
Metathorax bidentate: =. 2.2.2 2224. - (49) Amblyteles Wesmael. Metathorax umarmed.-..-.2:----.- (48) Pseudamblyteles Ashmead.
8. First abdominal segment at the elbow much swollen, gibbous, or angulated. (54) Probolus Wesmael. (Type, Ichneumon fossorius Gravenhorst. ) First abdominal segment at the elbow nof gibbous or angulated. Scuttellumepyramidale S22 222 eee! (55) Pyramidophorus Tischbein. (Type, Pyramidophorus flavoguttatus Tischbein. ) Scutellum not pyramidal. Antenne very strongly serrate. Male...(56) Pristocerus Grayenhorst. (Type, Pristocerus serrarius Gravenhorst. ) Antenne not strongly serrate. First abdominal segment neither broad nor rugose its entire Pere tise, eg ene ee set eh se ce es hanes hae 2) AG Ce he 9 First abdominal segment very broad and wholly rugose. (57) Rhyssolabus Kriechbaumer. (Type, Platymischos brassicus Tischbein. ) 9. Areolet pentagonal (rarely subtriangular and briefly petiolate in some males. ) Scutellum laterally margined at the most only at the base, never to the middle. Front tarsi without a single joint armed with fine spines. (58) Hurylabus Wesmael. (Type, Hurylabus torvus Wesmael. ) Front tarsi with most of the joints armed with fine spines. (59) EHristicus Wesmael. (Type, Ichnewmon clericus Gravenhorst. ) Scutellum laterally margined to beyond the middle. (60) Platylabus Wesmael. ) (Type, Platylabus rufus Wesmael. )
Tribe III. LISTRODROMINI.
1868. Listrodromoidx, Family 32, Forster, Verh. d. naturh. Ver. pr. Rheinl., XXV, pp. 144 and 194. 1894. Listrodromini, Tribe IV, AsHmEaAp, Proc. Ent. Soc. Wash., III, p. 278. The species belonging to this tribe have the claws pectinate, never simple; otherwise they are scarcely distinguishable from those found in the Joppini and the Ichnewmonini. Forster placed in the group only two genera, Veotypus and Listro- dromus, while I have ventured to place here five other genera,
TABLE OF GENERA.
Memtimmoracic spitactes- round Or Oval... 2.20. 2. seed sc Gees LeeLee Sec 4 Metathoracic spiracles, large, elongate, forming a long slit, sometimes curved. Scutellum flat, or at most subconvex, never gibbous or elevated _....-------- 2
Scutellum elevated at apex and highly declivous; metathorax normal, unarmed. (61) Cienochares Forster.
20 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXII.
Scutellum gibbous with lateral carinee; metathorax bidentate, exareolate. (62) Joppites Berthoumieu = Celmis Tosquinet = Pseudojoppa Kreichbaumer. Z0eMetathorax mornaal)) umn ed at ae See ee eae ee ar ee eee eee 3 Metathorax bispined or bidentate.
Spiracles of abdominal segments elongate or oval; metathorax not or very indistinctly areolated; submedian cell not longer than the median; disco-cubital neryure broken by a stump of a vein; areolet with the sides strongly convergent above, triangular or rhomboidal; abdomen banded, the ovipositor subexserted.
(63) Cressonianus Ashmead, new genus.
(Type, Patroclus lectus Cresson. )
3. Metathorax not short, not or very indistinctly areolated; submedian cell a little longer than the median; disco-cubital nervure broken by a stump of a
vein; areolet pentagonal; scutellum margined at sides anteriorly; abdo-
men blue or black, not banded, the spiracles of the first segment large, subreniform; claws with long teeth._-...---.-- (64) Patroclus Cresson.
(Type, Patroclus nigroceruleus Cresson. )
Metathorax short, truncate posteriorly and distinctly areolated; submedian cell a little shorter than the median, or never longer; disco-cubital neryure not broken by a stump of a vein; areolet regularly pentagonal; scutellum margined at sides clear to the apex; abdomen not wholly
blue or black, the spiracles of the first segment very small, rounded;
claws with shorter teeth at base only (sometimes difficult to discern).
(65) Neotypus Forster.
(Type, Ichneumon lepidator Fabricius. )
4. Metathoracic spiracles oval, the metanotum exareolated; scutellum flat, longer than wide, with elevated lateral margins.-..---.- (66) Hradha Cameron.
(Type, Eradha trichiosoma Cameron. )
Metathoracic spiracles round, the metanotum areolated; scutellum pyramidal. (67) Listrodromus Wesmael.
(Type, Ichneumon nyctermerus Gravenhorst. )
Tribe: LV." -HERESTARCE IMT: 1900. Heresiarchini, Tribe IV, ASHMEAD, Smith’s Insects of New Jersey, p. 567.
This tribe is proposed for ‘certain genera having the mandibles simple, edentate and acute at apex, and this simple character readily distinguishes the group from all others.
Four genera belong here, separable as follows:
TABLE OF GENERA.
Metathorax normal unarmed 22.2222 2.4. A= Se - s eee eeeeee 2 Metathorax bidentate.
Head large, strongly concave behind the temples, the cheeks full, buccate; transverse median nervure in front wings interstitial; disco-cubital nervure broken by a stump of a vein before the middle; antennz broadly ringed with white...---- (68) Plagiotrypes Ashmead, new genus.
(Type, Ichneumon concinnus Say.) 2. Metathorax with the areola semicircular, smooth, and shining; scutellum not margined laterally to beyond the middle; second abdominal segment with the gastrocceli linear and placed longitudinally. (69) Heresiarches Wesmael. (Type, Heresiarches eudoxius Wesmael.)
MRR oe
NO. 1206. ICHNEUMON FLIES—ASHMEAD. Di
Metathorax with the areola nof semicircular; scutellum: margined laterally to beyond the middle; second abdominal segment with the thyridia occu- pying the entire breadth and scarcely separated at the middle.
(70) Rhexidermus Forster.
(Type, unknown.)
Metathorax with the basal median and basal lateral areas confluent; scutellum
margined laterally only at base; second abdominal segment with the thyridia widely separated at the middle.
‘71) Stenodontus Berthoumieu. (=Gnathorys Wesmael. )
(Type, Ichneumon inarginellus Gravenhorst. )
iribe, Vo, ALOMY INI.
1844. Ichneumones heterogastri Wresmari, Nouv. Mém. Acad. Sci. Brux., X VITI, p-. 217.
1868. Alomyoide, Family 31, Forsrer, Verh. d. naturh. Ver. pr. Rheinl., XN V, pp. 144 and 194.
1894. Alomyini, Tribe III, AsHmeap, Proc. Ent. Soc. Wash., III, p. 278.
I have followed Forster in retaining this group as distinct from the Ichneumonini, where some authorities would place it, or from the Pheogenini, where others would place it. To me it seems to approach nearest to the Phwogenin/, but is readily separated by its metanotal characters and by the shape of the metathoracice spiracles.
Only a single genus is known in the group, distinguishable as follows:
Form elongate; metathorax smooth, exareolated, the spiracles large, oval; abdomen elongate, smooth and polished, the sides parallel, the second segment qwithout gastrocceli; antennze with the joints of the flagellum short, in female not or scarcely twice as long as wide, in male the joints, except the first, not longer than MUL MICA CIC MAGEALCC ys ears ona mee oS ei cke sclnceeee ss (72) Alomya Panzer.
(Type, Alomya ovata Panzer. )
drives Val PE ASTOGE NIN.
1868. Phxogenoidx, Family 30, Forster, Verh. d. naturh. Ver. pr. Rheinl., XX V, pp. 144 and 191.
1894. Phxogenini, Tribe V, AsHMEAD, Proc. Ent. Soc. Wash., III, p. 278
1898. Cyclopneustici, Subtribe, Berrnoumigu, Ann. Soc. Ent. France, L KV ps Oo2-
1900. Pheogenini, Tribe VI, AsuMEApD, Smith’s Insects of New Jersey, p. 568.
To this tribe belong a large number of the smaller ichneumonids, separated at once from those in the other tribes by the small, rounded, or circular metathoracic spiracles.
It is believed that the genera falling here can be readily distin- guished by the use of the following table:
TABLE OF GENERA,
Elica OLeSPeClallwAclevalteG@aemeceessccs sete ots te ese soe eee ecco eee 2
Scutellum very convex and elevated. Tip of abdomen acute, the ovipositor quite straight; head transverse-quadrate, the temples as wide as the eye; lunulze large transverse, more or less GCODMUCT teeta Oe eras aL Fe a (73) Ischnus Gravenhorst. (Type, Ischnus thoracicus Gravenhorst. )
bo
PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXII.
nw
Tip of abdomen very obtuse, the ovipositor curving upward. (74) Heterischnus Wesmael. (Type, Ichneumon pulex Miller.) 2. Superior hind angles of metathorax normal, not toothed..-..--...........-- 3 Superior hind angles of metathorax prominently toothed; clypeus subquadrate. (75) Apzleticus Wesmael. (Type, Apeleticus bellicosus Wesmael. ) Spiracles of first abdominal segment placed at the middle.
(76) Diacritus Forster.
Spiracles of first abdominal segment placed behind the middle.
Metathorax much lengthened and at apex produced beyond the base of the hind coxe. Clypeus convex, not separated from the face at base; abdomen not Gompressediaiihenpexe == aaa (77) Oronotus Wesmael. (Type, Oronotus coarctatus Wesmael.) Clypeus depressed, separated from the face by a deep furrow; abdomen compressed at apex -=2 2-21.33 (78) Diaschisaspis Forster. (Type, Diaschisaspis campoplegoides Holmgren.) 4. Second abdominal segment with the lunulze small, never twice as long as broad; metanotum nol sloping gradually from base to apex .....---.----- 5 Second abdominal segment with the lunulee very large, linear, twice as long as broad; metanotum gradually sloping from base to apex; areolet open’ of Gloseds es eae (79) Hemichneumon Wesmael. (Type, Hemichneumon suspectus Wesmael. ) 5. Areolet open behind; marginal cell along the costa scarcely longer than the triangular stigma; transverse median nervure in hind wings straight, NOU DRO KE Meteo ee (80) Epitomus Forster. (Type, Epitomus parvus Thomson. ) Areolet closed. Clypeustimarmed without ay tooth abapexees+ esse oe seat ene eee ee 6 Clypeus with a tooth at apex. Upper tooth of the mandibles longer than the lower; transverse median nervure in hind wings broken below the MMHG S.2 een co eee OEE eRe Ee Cee (81) Misetus Wesmael. (Type, Misetus oculatus Wesmael. ) 6. Clypeus at apex with a median semicircular emargination; mandibles with the teeth very unequalis 2s. esceee eee eee (82) Oiorhinus Wesmael. (Type, Oiorhinus pallipalpis Wesmael. ) Clypeus at apex without such an emargination. Clypeus at apex medially without a fovea - 4.2... .22212-- see sess sosebees G Clypeus at apex medially with a deep depression or fovea which often appears laterally as two small, blunt teeth; abdomen shagreened or densely coriaceous and finely punctate; mandibles rather large, the teeth subequal; transverse median nervure in hind wings broken very
little below the middle ..........------.-- (83) dithecerus Wesmael.
(Type, dthecerus dispar Wesmael. )
7. Discoidal transverse nervure wanting ....-.-.......-.... (84) Tycherus Forster. Discoidal transverse nervure present.
Second abdominal segment without distinct gastrocceli at base ...-------- 8
Second abdominal segment with distinct gastrocceli at base. Metathorax at apex not produced beyond insertion of hind coxe. Scape of antennze only slightly emarginate, longer than the first joint of flagellum ==s-- eos ee eee (85) Herpestomus Wesmael. (Type, Ichneumon brunnicornis Gravenhorst. )
j
NO. 1206. IGHNEUMON FLIES—ASHMEAD. es
Scape of antennee very deeply emarginate, shorter or no longer than
the first joint of flagellum. .--.----- (86) Diadromus Wesmael.
(Type, Ichneumon troglodytes Grayenhorst. )
Metathorax at apex produced somewhat beyond the insertion of hind COSKEE PEE pee tenn. LE eS See Oe ig (87) Thyrexella Holmgren.
(Type, Ischnus collaris Gravenhorst. )
8. Second segment with the thyridia more or less distinct -....2....--.------- 13
Second segment withowl a trace of thyridia or the same are unusually small and indistinct.
Mandibles in female at base beneath not emarginate ...--.------------- 9
Mandibles in female at base beneath emarginate. ; (88) Colpognathus Wesmael. (Type, Ichneumon celerator Gravenhorst. ) 9. Head quadrate or nearly, the temples broad Head transverse, not nearly quadrate. Metanotum with the areola lengthened, not cordate...---.---..\------ 10 Metanotum with the areola cordate or reniform. (89) Diceelotus Wesmael (—=Dicelus Wesmael=Cinwelotus Holmgren). (Type, Ichneumon pumilus Gravensorst. )
10. Scutellum margined laterally to the tip.-...----------- (90) Holocrepis Forster. Seutellum not marginal laterally to the tip; at the most margined only at the ; DELS Cnr ne eee ond on Oe ey ie (91) Deloglyplus Forster.
(Type, Deloglyptus punctiventris Thomsen. ) 11. Clypeus twice as wide as long; first abdominal segment somewhat longer than the second; flagellum in male very slender at base.......-.---- 12 Clypeus scarcely broader than long; first abdominal segment in female shorter than the second, in maleabout one-half as long. - (92) Micrope Forster. (Type, Pheogenes macilentus Wesmael. ) 12. Face much shortened; scape twice as long as the first joint of flagellum. (93) Eparces Forster. (Type, Eparces quadriceps Thomson. ) Face not much shortened; scape stout, globose. First joint of flagellum rarely longer than thick, shorter than the second; transverse median nervure in hind wings broken below the middle. (94) Centeterus Wesmael. (Type, Centeterus major Wesmael. ) First joint of flagellum elongate, three or more times longer than thick, and longer than the second; transverse median neryvure in hind wings broken above the middle.-..---------- (95) Pecilostictus Ratzeburg. 13. Mesonotum and: scutellum not wholly flattened -.........--.--------.----- 14 Mesonotum and scutellum wholly flattened, the postscutellum smooth, shining. Metanotum with a distinetly circumscribed areola----. (96) Mriplatys Forster. (Type, Harpestomus ardeicollis Wesmael. ) Metanotum without an areola -......----.----- ...-(97) Anopiesta Forster. 14. Second abdominal segment with two fovezw at base; metathorax not areolated. (98) Nematomicrus Wesmael. Second abdominal segment with thyridia only at base; metathorax areolated. Thyridia lying close to the base and indistinct; postpetiole broad, strongly punctured; clypeus thickly punctured. ----- (99) Beosemus Forster. (Type, Ichneumon mitigosus Gravenhorst. ) Thyridia not lying close to the base and usually large; postpetiole not broad nor strongly punctured; clypeus not thickly punctured. Postpetiole very short, scarcely one-fourth the length of the petiole;
24 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXIII.
thyridia very large and broad, placed far beyond the base and only a little before the middle of the segment.
Head quadrate; areola pentgonal or nearly; abdomen sha-
greened or punctate basally. -....- (100) Notosemus Forster.
(Type, Notosemus Bohemani Wesmael. )
Head subglobose; areola of metanotum semicircular; abdomen
SMmOOth= ees Fe ee eee ee (101) Mavesia Holmgren.
(Type, Phogenes argutus Wesmael. )
Head not quadrate, at most subquadrate; postpetiole not very short;
thyridia placed tolerably close to the base of the segment; clypeus
completely separated from the face.
Cheeks not buccate; the elypeus truncate or slightly rounded
antenorhytes 5 7\25= ao eee (102) Phxogenes Wesmael.
(Type, Ichneumon semivulpinus Gravenhorst. )
Cheeks buccate; the clypeus very short bisinuate anteriorly.
(103) Proscus Holmgren.
(Type, Phxogenes cephalotes Wesmael. )
Subfamily Il. CRY PTIN 4.
1868. Cryptoidx, Family 26, F6rster (part), Verh. d. naturh. Ver. pr. Rheinl., XXV, pp. 144 and 186.
1873. Cryptidx, Family, THomson, Opus Ent., V, p. 467.
1887. Cryptine, Subfamily, Cresson, Syn. Hym. North America, p. 42.
1888. Cryptidx, Family, THomson, Opus Ent., XII, p. 1286.
1900. Cryptine, Subfamily II, AsHMEap, Smith’s Insects of New Jersey, p. 568.
This subfamily, with the exception of possibly the /chnewmonina, ‘an be easily separated from all the others by the characters made use of in my table of subfamilies. With the subfamily mentioned, however, it is different, since the species falling in it are exceedingly closely allied, and the males especially are separated, or placed, with difficulty. The females, however, may be easily distinguished, “except in a few cases, by the prominent, exserted ovipositor, and the position of the spiracles of the first abdominal segment.
Both sexes, however, possess a character not found in the /ehneu- moninne, Viz, «more or less distinct, longitudinal grooved line or furrow, sometimes punctate or crenulate, situated low down on the mesopleura and which separates the mesosternum from these sclerites. This char- acter may always be depended upon to separate a cryptine from an ichneumonine.
Seven distinct minor groups, or tribes, may be distinguished, sep- arated as follows:
TABLE OF TRIBES,
Metathorax without distinct longitudinal carinee or at the most with only the pleural carinze present, the petiolar area always wanting, usually with one or two transverse carinze or with none; stigma most frequently nar- rowed, lanceolate; apterous and subapterous forms common. .-.-- a
Metathorax with longitudinal carinee and usually more or less completely areolated, the petiolar area present; stigma usually widened, triangular, sub- triangular, or ovate; subapterous forms rare.
pin
No. 1206. ICHNEUMON FLIES—ASHMEAD. 25
Metathorax usually produced beyond the insertion of hind cox, the petiolar area and the areola usually confluent and extending clear to the base; ovipositor very short, at the most subexserted.
Tribe I. Srrupnint.
Metathorax not produced beyond the hind coxse; ovipositor exserted.
Front wings with a complete areolet; head usually quadrate; antennze and legs most frequently stout -...-------- Tribe Il. PryGaprvontint.
Front wings with the areolet incomplete, open behind or wanting; head not or rarely quadrate; antennze and legs usually slender.
Basal nervure not strongly curved inwardly...-Tribe ILL. Hemrrenrnt.
Basal nervure strongly curved inwardly --.---- Tribe TV. Przomacnint.
PAROS SBhUlllveCevelLOMe daa aaa Ae on ERO LSE te eatin 3 Wanes albsemtronainlneviatedss ss50 aa 2s feet eS Beate Tribe IV. Prezomacnint. Metanotmm motjareolated(.s {325.02 gb 2 ale e2 Tribe IV. Przomacurnt.
3. Front wings with the stigma narrowed, the areolet variable, pentagonal, or small quadrate, sometimes almost punctiform, more rarely open behind or entirely absent; discoidal cell with the lower apical angle straight or obtuse, the basal nervure not strongly curved inwardly; ADS CiSSsanOimCOstailOM ey =e tea et ee ken sl 2 den oe Se cigs 4
Front wings with the stigma broad, triangular, the areolet pentagonal in posi- tion but open behind, the basal nervure strongly curved inwardly;
aloscissaxshort: (Viale?) seas 28 ee ee eee Tribe IV. Przomacnuint. MeeAreoOletmentireliy, wamtines.2.+.----4-.-225-255------ Tribe V. Hemicasrrrint.
Areolet distinct, usually large, never very small, and always pentagonal, the sides convergent above or parallel. -..-------- Tribe VI. Cryprint. Areolet small, quadrate, sometimes almost punctiform, sometimes open behind, but never pentagonal in position... ~~~ --- Tribe VII. Masosrentnt.
‘ribo owk Selec PNINE
1868. Stilpnoide, Family 28, Forsrrr, Verh. d. naturh. Ver. pr. Rheinl., X XV, pp. 144 and 188.
1873. Stilpnina, Tribes, Tomson, Opus Ent., V, p. 468.
1884. Stilpnina, Tribes, THomson, Opus Ent., X, p. 1018.
1894. Stilpnini, Tribe 1, AsHmran, Proc. Ent. Soc. Wash., III, p. 278.
1900. Stilpnini, Tribe I, ASHMEAD, Smith’s Insects of New Jersey, p. 568.
This group is of small extent and at one time, on account of the brevity of the ovipositor, was confused and classified with the genuine Ichneumonines; but from these it is at once separated by the longitu- ninal furrow which separates the mesosternum from the mesopleura.
The tribe is distinguished from the others in this subfamily not only by the non-exserted, or at most, subexserted ovipositor, but also by its metathoracic characters. All, except two or three of the genera, have the metathorax long, gradually sloping from base to apex, and pro- duced posteriorly beyond the insertion of the hind coxe, with the areola and the petiolar area confluent, extending to, or almost to, its base.
Most of the species, too, are highly polished and have the abdomen long, more or less compressed, rarely short or broad, while the areo- let in the front wings, although sometimes closed and pentagonal, is most frequently wanting or open.
26 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL, XXIII.
Kight genera are placed here, distinguished as follows: TABLE OF GENERA.
Fourth abdominal segment and the following not at all or only slightly compressed; if much compressed, compressed from the second segment, the incisions
always distinctly: visible: .~ 222552. <4. se te So eee ee 2
Fourth abdominal segment and those following very strongly compressed, the IMGISIONS) scarcely visio lesa = === ane (104) Selewcus Holmgren.
(Type, Seleucus cuneiformis Holmgren.)
2. Third joint of flagellum strongly excised.-..........--- (105) Zetisima Forster.
(Type, Zetisima rufipes Forster. ) Third joint of flagellum not excised.
Areolet closed at apex, or, if open, the abdomen much lengthened _.__-- 3 Areolet open at apex, the abdomen rounded or oval; antennze 17-18-jointed. (106) NXestophya Forster. (Type, Nestophya fallax Forster. ) 3. Second abdominal segment in female from the base and beyond not much com- pressed, the postpetiole not entirely smooth and shining -.......__-- 4 Second abdominal segment from the base and all the following segments much compressed from the sides, the petiole entirely smooth, shining, the postpetiole scarcely wider than the petiole; second segment longer than wide at apex; metanotum with the external and median lateral areas Goutlnemt 2% ee ees ae at ee aha ce eo eae (107) Asyncrita Forster. (Type, Atractodes foreolatus Gravenhorst. ) 4. Antenne in female 16-17-jointed, in male 19-23-jointed; pronotum anteriorly TUN COVE red eee See = at Ae ota ee gee (108) Stiépnus Gravenhorst. (Type, Stilpnus gagates Gravenhorst. )
Antenne in female more than 17-jointed; pronotum covered.
Abdomen in female either lengthened or somewhat compressed from the sides, the second segment more or less rounded laterally; areolet either closed or open behind
Abdomen in female compressed laterally, also not strongly lengthened, the second segment laterally not rounded, much widened toward apex; spiracles of the first and second segments in males and females not really visible from above: .---.-.---------- (109) Polyrhembia Forster.
5. Abdomen in female not compressed laterally, with a distinct ventral fold; mid- dle vein in hind wings obliterated at base in both sexes; petiole and post- petiole in male smooth, shining, and longer than the coxe and trochan- ters; second segment with thyridia; areolet open.
(110) Exolytus Holmgren. |
(Type, Mesoleptus levigator Grayenhorst. )
Abdomen in female much compressed laterally, without a ventral fold; middle vein in hind wings distinct in both sexes; petiole in male more or less coriaceous or rugulose, not longer than the coxze and trochanters; areolet WarltingOriopens Aso: © eee apa eee: (111) Atractodes Gravenhorst.
(Type, Atractodes bicolor Gravenhorst. )
aN kN al lt Nl a lA aie aaa a Al tlt alld een at A Slt
Ser eres ami
itt cake ed! BONE,
4 ie
NO. 1206. ICHNEUMON FLIES—ASHMEAD.
t |
iicicoey ale I ri rGADEUONINI:
1868. Phygadeuontoide, Family 25, Forsrer, Verh. d. naturh. Ver. pr. Rheinl., XXV, pp. 144 and 181.
1873. Phygadeuonina, Tomson, Opus. Ent., V., pp. 468 and 517.
1894. Phygadeuonini, Tribe III, AsumEap, Proc. Ent. Soc. Wash., III, p. 278.
1900. Phygadeuonini, Tribe I], ASumMeEap, Smith’s Insects of New Jersey, p. 568.
This group is undoubtedly the largest and probably the most dif_i- cult to study and classify of all the tribes in this subfamily, and I am by no means satisfied that, as at present constituted, it is a natural group, or that it can always be kept separate from the //emitelini. In fact, 1am inclined to believe, from studies thus far made in the two groups, that some of the forms now placed in the //emiteliné really belong here, and that the closed or open areolet will not always prove a reliable character to separate them, although typical forms are readily separated by it.
For the present, however, or until a larger and better collection can be brought together, I prefer to treat these two tribes in the Fors- terian sense.
About 51 genera are now recognized in the tribe, separable as follows:
TABLE OF GENERA.
Bees mOrinial wn ObabNreviatede sss a2 2 kes eo see se oes eee ones 3 Wings abbreviated. Mainoseiironcl osedanmancinalecellizee ees hae eee 8 2 ye et Sale 2
Wings without a marginal cell.
Basal nervure present; metanotum incompletely areolated, the apical trans- verse carina distinct, complete-----.---- (112) Stibeutes Forster. (Type, Stibeutes gravenhorstii Forster.) Basal nervure absent; metanotum not areolated, the apical transverse carina AC OMp LELORPee ees Sates =e ye See ee (118) Pezoporus Forster. (Type, Pezomachus nigrocinctus Gravenhorst. )
2. Wings with a basal nervure; metathorax completely areolated. (114) Phyrtus Forster. (Type, Hemiteles hemipterus Gravenhorst. ) Wanes without basal neryure-.....-.¢..--.<-2.--<- (115) Chamezelus Forster. 3. First three joints of flagellum not especially lengthened, or at least not the third; in female scarcely more than twice, or at most thrice, as long as thick; in male rarely more than thrice as long as thick at IDEA eet ne ae Weer At ae See AM oe eae So LS YR 8 First three joints of flagellum much lengthened; the first and second at least four times as long as wide at apex, or longer, the third fully three or more times longer than thick ......+.......--.---- 4 4. Sides of the face clothed with a glittering or silvery white pubescence; parap- sidal furrows deep and extending at least to the middle of the VE SON LUM aera a seen eae teres ts eye Sy se 2 Seo 5 Sides of the face not clothed with a glittering white pubescence; parapsidal furrows wanting or indistinct, at the most feebly indicated ANCOR OL yee meee ea een sae = ei a aclS cis clavelc.cicicin a.cie' «= 6
ad
28 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL, XXIII.
5. Transverse median nervure in hind wings not broken. (116) Thysiolorus Forster. Transverse median nervure in hind wings broken. Disco-cubital nervure straight or slightly curved, but not angularly broken. Abdomen scarcely longer than the head and thorax united, the second segment at apex thrice as wide, or nearly, as long. (117) Apsilops Forster. Abdomen longer than the head and thorax united, the second segment longer than wide at apex; areolet with the sides convergent above. Metathoracic spiracles large, long elliptical (male) (see p. 29). (126) Plectocryptus Thomson. Metathoracic spiracles small, short oval or subrotund (male) (see TOS ZO) rs ra ae ee te (127) Microcryptus Thomson. Disco-cubital nervure angularly broken. Abdomen elongate, much longer than the head and thorax united, the second segment not twice as wide as long, not much more than
half the length of the segment. ~~... - (118) Panargyrops Forster. 6. Metathorax regularly areolated, more or less rugolose, or coriaceous, and fre- quenthy opaque ++... 2.2 2 a ee ee ee ee 7
Metathorax areolated, but quite smooth and shining.
First abdominal segment with dorsal carinve; metathorax with jive areas at BPO Ae see ee See tes Bene eee (119) Leptodermas Forster. First abdominal segment without dorsal carinze; metathorax with thrce areas ait BESET Se Fee meee so eer en te oe (120) Oxytenia Forster. Radius originating before the middle of the stigma; disco-cubital cell at base as wide as the second discoidal cell at apex. -(121) Isotima Forster. Radius originating from the middle of the stigma; disco-cubital cell at base
nearly twice as wide as the second discoidal cell at apex. Transverse median neryure in hind wings obtusely angularly broken a little above the middle; petiole long, almost straight, not elbowed or much widened at apex.......-... (122) Acroricnus Ratzeburg. Transverse median neryure in hind wings straight and broken by the sub- discoidal nervure far below themiddle; petiole bentand widened GTO au MGT Te pees Oe eee ey eee eee (123) Stiboscopus Forster. 8. Dorsal carinze of first abdominal segment extend from the base to the spiracles, but motibeyondes. 2222222 psa a asec ee eo 9 Dorsal carinze of first abdominal segment extend from the base to beyond the spiracles, but rarely to the tip of the segment; if not, then antennze in female compressed or flattened between the middle and thesapexd see Se (See See ei eS ae 10
=I
<2
Hind tibize deeply incised at apex, the tarsi attached below the tip. Metanotum areolated, the areola wider than long; hind tibize spinulose. (124) Glyphicnemis Forster. (Type, Phygadeuon vagabundus Gravenhorst. ) Hind tibize normal, not deeply incised at apex; the tarsi attached normally. Metanotum with the lateral basal and median areas not confluent. Spiracles!smal lero una ee eee (125) Bathymetis Forster. Spiracles long oval or ovate (males). Last joint of tarsi as long as the third; scutellum spotted with
yellows (see po 2i)i.ises. ahs eee (113) Pezoporus Forster. Last joint of tarsi shorter than the third; scutellum black (see Dizi) eosaseee ees 5. Hote ee (112) Stibeutes Forster.
NO. 1206. ICHNEUMON FLIES—ASHMEAD. 29
Metanotum with the lateral basal and the median areas confluent. Spiracles rather large, elliptic-oval -....-- (126) Plectocryptus Thomson. Spiracles rather small, short oval or subrotund. (127) Microcryptus Thomson. (Type, Cryptus erythrinus Grayenhorst. ) 10. Clypeus in male and female anteriorly distinctly bidentate, or with two, more
OrALES SCS GIN Ciaplnl) PLES meee ames teen See emiss ose se 11 Clypeus with the anterior margin simple or with a single tooth.._....----- 13
yt ( g ; MimeLiwer pare. meverdisnmenly hairy s.25- 2. .-22--52s-5 6 ec2s2-25 5.222545 eee. 2
Eyes distinctly hairy. Antenne tricolored, ringed with white; first and second flagellar joints of
aMvequalelenet haces «sarees ees Sree (128) Iselia Forster.
Antenne neither tricolored nor ringed with white; first flagellar joint shontembhamtme:second.-2-22--—-+-.2-- (129) Homelys Forster.
12. Metanotum at base not completely areolated......---- (130) Polytribax Forster.
Metanotum at base completely areolated.
Jarina at apex of the middle lateral area sharply elevated; second segment much narrowed toward the base, scarcely half as wide as at apex, and finely striately rugulose its entire length.
(181) Ernoctona Forster. Carina at apex of the middle lateral area not sharply elevated; second seg- ment not much narrowed toward base, more than half as wide as at apex, and not striate its entire length. (182) Plesiognathus Forster. (Type, Phygadeuon cephalotes Gravenhorst. ) 13. Clypeus with one tooth on its anterior margin. --- - (183) Micromonodon Forster. Clypeus with the anterior margin simple or without a tooth.
Transverse median nervure in hind wings not broken, or broken below the Ta CLC G pene) a ere ch oR aterm eie Tat Seen eee toate 14
Transverse median nervure in hind wings broken aé or above the middle. Transverse median neryure in front wings originating before the basal neryure; base of third discoidal cell much wider than the base of the second discoidal cell.....-.--- (134) FHeterotypus Forster. Transverse median nervure in front wings nof originating before the basal nervure; base of third discoidal celi not wider than the base of the second discoidal cell.__-.--- (135) Dapanus Forster. = Sorbas Forster = Trichocryptus Forster. (Type, Ichneumon cinctorius Fabricius. )
14. Transverse median nervure in hind wing broken below the middle. --..-.-- 15 Transverse median nervure in hind wings wot broken.
Abdominal segments 2 and 3 very large..----------- (186) Hedylus Forster.
15. Pronotum not lengthened; ovipositor prominently projecting. ......-------- 16
Pronotum lengthened; ovipositor only slightly visible beyond the tip of the DOGO Me Tepe tegen cern ce aN tee ee (187) Dirophanes Forster. 16. Petiolar area very short, the areola narrow, rectangular, extending to apex; head very small; antennze slender, filiform. (188) Tricholinum Forster. Petiolar area not very short, the areola not long, rectangular, most frequently transverse and hexagonal, rarely pentagonal, if elongate, nar- rowed toward base, rarely wholly wanting.
Kyes hairy. Second abdominal segment shorter than the third......--.------- 17
30
20.
21.
sy)
PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXIII.
Second abdominal segment a little longer than the third, smooth and polished, the post petrole striate. ------ (139) Zaphleges Forster. (Type, Phygadeuon leucostigimus Gravenhorst. ) Foyea at base of scutellum divided by a sharp carina, metanotum completely areolated, the areola transverse, trapezoidal. (140) Endasys Forster. Fovea at base of scutellum wot divided by a sharp carina; metanotum completely areolated, the areola longer than wide, hexagonal. (141) Baryntica Forster. Middle joints of flagellum above in female not flattened, in male clothed usually with. short, :shagey- hairs. - 22. 22ee Bees Soe te Oe eee 19 Middle joints of flagellum above much flattened. Metanotum exareolated or very incompletely areolated; spiracles large, linear orellipticalive. s+ 22.5 +e eee (142) Giraudia Forster. (Type, Cryptus congruen Gravenhorst. ) Metanotum with a long middle area, the areola and basal area very united; spiracles not large, oval; subdiscoidal nervure in hind wings originating very close to the origin of the transverse median Eo ee area ae Nereida (143) Schenkia Forster. (Type, Cryptus graminicola Gravenhorst. ) Metathorax with four distinct prominent teeth; disco-cubital neryure broken by a stump of avein near the middle - (144) Rhembobius Forster. (Type, Phygadeuon quadrispinosus Gravenhorst. ) Metathorax at most with two prominent teeth, often unarmed. Hind stibice:mormaill sot 26s fae One ed oe Sa rel 20 Hind tibize toward apex broadened and broadly flatly truncate. (145) Colocnema Forster. Metanotum at base usually more or less incompletely areolated, the areola and basal areas confluent, or the former is not separated from the
middle lateral areas by a sharp carina: _< 2222-22202 22-528 21 Metanotum) atibaseicomipletelyaneolated eae 5 = ee ee 24
Lower tooth of mandibles much longer than the upper tooth. Head quadrate; transverse median neryure interstitial with the basal nery- UPO% cs Sodas a ee ea Ree (146) Ecporthetor Forster. (Type, Phygadeuon fortipes Gravenhorst. ) Lower tooth of mandibles shorter or no longer than the upper tooth. Metathoracic spiracles round, or very short oval, scarcely longer than Wide (So eae 52 Ps ae ee ean a ay ge Se 22 Metathoracic spiracles fully twice as long as wide, or nearly. Areola seen from aboye pyramidal; metathoracic spiracles not quite
iwiceas lonclasnmidesss-— eee] =e (147) Neleophron Forster. Areola seen from above not pyramidal; metathoracic spiracles twice or more than twice longer than wide..-.(148) Epiphobus Forster.
Head cubiform. ; Femora somewhat short and swollen; antennze short, stout; metanotum without or with areas confluent -----.- (149) Ecpaglus Forster.
(Type, Phygadeuon brevicornis Gravenhorst. ) Head not cubiform.
Disco-cubital nervure with a short process. ----- (150) Odontoneura Forster. Disco-cubital nervure without a process. Second recurrent neryure received by the areolet at or behind the mid- dle 2.22 32354232 ee eee 23 Second recurrent nervure received by the areolet before the middle. Metanotum coarsely rugose, the areola very high and narrow; first abdominal segment wholly striate...(151) Ulothymus Forster.
i :
No. 1206. ICHNEUMON FLIES—ASHMEAD. ak
Metanotum not coarsely rugose, the first and second lateral areas
confluent; first abdominal segment not striate. (152) Ophidnus Forster.
23. Lower tooth of mandibles very small and much shorter than the upper tooth. (153) Homotherus Forster. Lower tooth of mandibles equal, or nearly, with the upper tooth. Posterior tibize and tarsi normal, not spinulose.
First three segments of abdomen finely coriaceous, the second a little longer than the third....---.------ (154) Pammachus Forster. = Stenocryptus Thomson. First three segments of abdomen smooth, the second and third of an
equalglengtihgee sees sense ase (155) Phygadeuon Gravenhorst.
Posterior tibize and tarsi spinulose. iMetathoraxabidentates=s2-- 224-2 4- 552 (156) Trachyphyrus Haliday. 24. Spiracles of the second and third abdominal segments placed close to the lateral TUNG OU een es eo aoe ta ee me eam ae che Ste Bet torte he 25 Spiracles of the second and third abdominal segments placed away from the PAGer el enna DOAN ot cere Mee ee Oe Grats ens ames ts. Aaa a rR the 26 PyeScute lumEVery Wat, oo coe se ee eos se chen eto ae (157) Terpiphora Forster. Seutelilmmrconvexa- pean ee ak eeee. & see's tyne (158) Scinascopus Forster. 26. Third abdominal segment not longer than the second._..........-.-.------ ai Third abdominal segment longer than the second... (159) Medophron Forster. 27. Metanotum with the areola most frequently hexagonal, never pentagonal, the basaltaneaimeverstmanculare. = 42-022 sl2- a... 2222s see 28 Metanotum with the areola regularly pentagonal, quite pointed toward apex, the basal area triangular -...---------- (160) Phyzelus Forster. 28. Anterior margin of clypeus not emarginate; metathoracie spiracles surrounded by Sharp curved) caring: ---.2252222-2-5: (161) Bachia Forster.
Anterior margin of clypeus more or less emarginate. Metanotal carina angular; radius originating from the middle of the stigma; first abdominal segment with strong dorsal carine.
(162) Nuneches Forster. Metanotal carina curved; radius originating behind the middle of the stigma; first abdominal segment without dorsal carinze; upper
tooth of mandibles more than twice as long as the lower. (163) Demopheles Forster.
perioer bie = CE Mire TEIN I:
1868. Hemiteloide, Family 24, Forster, Verh. d. naturh. Ver. pr. Rheinl., X XV, pp. 144 and 173.
1873. Hemitelina, Tribus, THomson, (part) Opus. Ent., V, p. 468.
1884. Opus Ent., X, p. 967.
1894. Hemitelini, Tribe II, ASHMEAD, Proc. Ent. Soc. Wash., III, p. 278.
1900. Hemitelini, Tribe II], AsHmeap, Smith’s Insects of New Jersey, p. 569.
Thomson (see above) included with this tribe Férster’s Pezoma- choide, but so far I have been able to separate the two readily by the characters made use of in my table of tribes, the female being distinguished by metathoracic characters and the winged males by the difference in the shape of the basal nervure in the front wings.
Forster gave no character to separate the winged males in this group from those in the Pezomachini, and I suspect he may have included
32 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXIII.
some of them here under different generic names, since I have already recognized three or four generic types of malesamong the Pezomachini.
Some 78 genera fall in this tribe, as at present interpreted, although some of these, if I have identified them correctly, will have to be removed to other tribes later, that is, to the Phygadeuonini Pezomachini, and possibly to the Plectiscind.
TABLE OF GENERA.
First transverse cubitus not entirely wanting, usually very distinct, the areolet pentag- onal in position, but open behind, the transverse neryure entirely wanting or very pale, subobsolete.---...---.------- 3 First transverse cubitus entirely wanting, the discocubital nervure being interstitial with the second abscissa of the radius, the areolet wholly wanting
« ~
2. First joint of flagellum as long or somewhat longer than the second; vertex as high as the upper eye margins. Ocelli lyanevclose towhereyes=s2= 255 sae soe (164) Spinolia Forster. Ocelli not lying close to the eyes, the lateral ocelli as wide, or nearly, from each other as to the eye margin. Antennee 20-jointed or more _.....-..-..------- (165) Allocota Forster. Antenne short, less than 20-jointed._(166) A/lastoneura Kriechbaumer. First joint of flagellum shorter than the second; vertex much higher than the upper eye margins; ocelli far away from the eyes; eyes small; anitennce sl /(-]olnbe ds (167) Syneches Forster. oe vetanofummn ot atrallvarecolatedssssees: haere eee eee (168) Chirotica Forster. Metanotum more or less areolated. Second discoidal cell closed SecondidiscordaltcelllRo werk aith ct exae eases eee eee 4 4. Wings in female much shortened, without a stigma; head quadrate. (169) Catolytus Forster. Wings normal, with a stigma; head transverse. Antenne 15-17-jointed; metathorax with the petiolar area very large. (170) Gnypetomorpha Forster. Antenne at least 19-jointed; metathorax regularly areolated. (171) Nenolytus Forster.
b: Metathoracie'spiracles ‘round 62 2st ee a ee ee ee 6 Metat horace: spiracles yell geese eee ee (172) Otacustes Forster. 6. Transverse median nervure in hind wings broken.............------------ 15
Transverse median neryure in hind wings not broken, straight. Antennze more than 17-jointed; second abscissa of the radius not 5 times as lons-as-the first i222 sec bs so Se See ee ies eee a Antenne 17-jointed; marginal cell very long and pointed, the second abscissa of radius about 5 times as long as the first.
(1738) Cenomeris Forster.
7:-Discoidal: cell:closedat apexes .2a22 soce eee ee ee eee 8 Discoidal cell open at apex. Second discoidal cell closed.-.-.._.-.-.--.--------- (174) Acrolyta Forster. 8 blindstemoranverys tiicke 55 ys. se aes eee (175) Gunopaches Forster. Hind femora not very thick, normal. Metanotum with the carina distinct, not obliterated at the middle -...-- 9
Metanotum with the carina obliterated at the middle. Parapsidal furrows uniting at the middle of the mesonotum; areolet irregular; first abscissa of the radius fully half the
sa cat ah ee
}
| 2 an
% ICHNEUMON FLIES—ASHMEAD. 33
length of the second; first joint of the flagellum longer than the
a Be COU Ue eee ey ee eae tye ae (176) Thestis Forster. 3 Parapsidal furrows not uniting at the middle of the mesonotum; h areolet regularly formed: first abscissca of radius not half the * length of the second; first joint of flagellum shorter than the ' RCE Se meen Rr re ered Ph ae Ne AS (177) Pemon Forster. 9. Metathorax with the petiolar area bounded by a prominent ridge above, the middle lateral area also prominent —__—-- (178) Trisacra Forster.
Metathorax with the petiolar area not bounded by a prominent ridge above. Metanorummcounpietely vareolated :)2 202. 2. 22k ese tek I4
Metanotum completely areolated. Face not clothed with long glittering white hairs; mesonotum with the parapsidal furrows incomplete or wanting......------- 10 Face clothed with long glittering white hairs; mesonotum with the parapsidal furrows complete.
Face very much narrowed -.------ (179) Ischnurgops Forster. ACEH TOA ee errs ee ne eel eee (180) Steganops Forster. 10. Middle lateral areas very strongly toothed_........---- (181) Ischyracis Foster.
Middle lateral areas not strongly toothed. Clypeus distinctly separated; anal valves in male small; mandibles not emarcinateatthe middless= os enn) De eae es Se 11 Clypeus not separated, wholly bent downward, the anterior margin squarely truncate; anal valves in male very large, prominent; mandibles very small, emarginate medially. (182) Astomaspis Forster. iMpeletiolar area not confluent with the areola-~....-..2.------------5-+++---- 12 Petiolar area confluent with the areola and extending nearly to the base of the metanotum; antenne 18-jointed, toward apex clavate. (183) Microplex Forster. 12. First joint of flagellum fully as long or longer than the second.......------ 15 First joint of flagellum a little shorter than the second. ...(184) Lysibia Forster. 13. Middle vein in hind wings toward the base obliterated and only visible by a EN eublnne MM Crier rar a eee lene eye e cine om sis (185) Daétora Forster. Middle vein in hind wings distinct, not obliterated at base. Metanotum with 5 areas; spiracles of the first abdominal segment very
JORRO UMA Pe ae Oia e es ReneS (186) Aclastus Forster.
Metanotum with 3 areas; spiracles of the first abdominal segment not at all OOM E TILE = Sats sere cel om eee <a oe (187) Opisthostenus Forster.
14. Metanotum without areas at base; wings very narrow. (188) Asthenoptera Forster. Metanotum with areas at base; wings broad _...--------- (189) Stygera Forster.
15. Metathoracic ridge not interrupted at the middle..-............-+--------- 16
Metathoracic ridge interrupted at the middle. First abominal segment short, broad, and strong --(190) Diaglypta Forster.
Memes Gishinctlyhalrys. 2.2 200s 2 fa seso so ee ee (191) Habromma Forster. Eyes not hairy. Ocellisiouching the eyes:-=.2-.--...-- .----+--44=- (192) Plesiomma Forster. Ocelli not touching the eyes. Clypeus bare, or nearly, without long hairs ........----------- 17
Clypeus with long hairs, almost forming a fascicle. (193) Bathythrix Forster.
17. First joint of the flagellum somewhat shorter than the second -..-.-.------- 18 First joint of the flagellum as long as or longer than the second. -.---------- 19
18. Third joint of the flagellum as well as the second joint somewhat longer than the first; clypeus anteriorly at the middle impressed and broadly,
although slightly, emarginate..---------- (194) Algina Forster. Proc. N. M. vol. xxiii——3
o4
1g.
20.
24.
20.
26.
oT
PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXII. |
Third joint of the flagellum not longer than the first; ¢lypeus anteriorly at the middle neither impressed nor emarginate.(195) Daictes Forster. Transverse median nervure in hind wings broken at or below the middle, never above the middle: so: vs. esky CN eee ora ieee soe nee pene 20 Transverse median neryure in hind wings broken above the middle. (196) Strepsimallus Forster. Head widened behind the eyes, the temples broad---- - (197) -Enoplex Forster. Head not widened behind the eyes, the temples flat or narrow. Discoidal nervure not longer than the base of the discocubital cell Discoidal nervure longer than the base of the discocubital cell. (198) Mastrus Forster. Metanotum with the apical carina strongly angulated or toothed. (199) Lymeon Forster. Metanotum with the apical carina normal. Radius more or less curved, but not broken at a right angle..._......-- 22 Radius broken almost at a right angle -__..._.-- (200) Paraphylax Forster. Discoidal cell not narrowed at base, the apex lying far from the hind margin of the’ Wines - LA see ee eee bee eee te eee ee 23 Discoidal cell much narrowed at base, the apex not far from the hind margin of wings, the entire radius strongly arcuate; transverse median nervure in hind wings quite near the inner margin. (201) Rhadiurgus Forster.
Metanotumyuhouticamnines === =e eee ee (202) Aschistus Forster. Metanotum with carine. Clypeus anteriorly without an impression _............2.2--.-2.----+-- 24
Clypeus anteriorly with an impression on both sides. Metanotum coarsely rugose; antennze in male thick, the first joint of the flagellum not thrice as long as thick--__. (203) Tolmerus Forster. Metanotum not coarsely rugose; antennze slender, filiform, the first three joints of flagellum at least five times as long as thick. . (204) Rhadinocera Forster. Clypeus anteriorly not bidentate; middle lateral areas not broadly carinately
prominent atapex 22: = as Ser oe Sate ee ee 25 Clypeus anteriorly at the middle bidentate; middle lateral areas at apex broadly carimnately prominent. s2nse= ss sa see (205) Isadelphus Forster.
Penultimate joint of the maxillary palpi more than half as long as the last.. 26 Penultimate joint of the maxillary palpi only half as long as the last. (206) Blapsidotes Forster.
Third joint of hind tarsi as long as or longer than the fifth -..........2-2-- 27 Third joint of hind tarsi shorter than the fifth.
Spiracles of the first abdominal segment placed somewhat before the mid- dle; ovipositor with a slight upward curye.
(207) Allomacrus Forster. Third joint of hind tarsi longer than the with..: = seosse2 eee 29 Third joint of hind tarsi of an equal length with the fifth.
Clypeus distinctly but not deeply separated; all femora, and especially the hind pair, distinctly thickened; head much narrowed behina the eyes; the middle lateral areas at apex not strongly promi- Nent ccs X = Jeb ek ee eae HOS eee er ae 28
Clypeus very deeply separated; femora not especially thickened; head not especially narrowed back of eyes; metathorax with five areas at apex, the middle lateral area strongly prominent at apex.
(208) Philonygmus Forster.
dnt an tel ll
so inl te sic
NO. 1206. ICHNEUMON FLIES—ASHMEA D. 35
28. Metathorax at apex perpendicularly truncate, the carinze not sharp, the petiolar and lateral apical areas confluent; first abdominal segment at apex twice as wide as at base; disco-cubital nervure broken by a stump of a vein; antennze usually ringed with white.
(209) Barydotira Forster.
Metathorax at apex not perpendicularly truncate, the carinze very sharp, the petiolar area separated from the lateral apical areas; first
_ abdominal segment at apex not much wider than at base;
antennz not ringed with white _____- (210) Pantolispa Forster. 29. Metathorax as seen from the side perpendicularly truncate, or almost-.----- 30 Metathorax as seen from the side not perpendicularly truncate._.....--.__- Bil
30. Dorsal carine of the first abdominal segment sharp and distinct from the base to beyond the middle; first joint of the flagellum scarcely longer than the second, the latter distinctly longer than the third, the ten joints before the last in female wider than long; marginal cell not longer than the stigma -...--- (211) Microtorus Forster.
Dorsal carinze of first abdominal segment extending to the middle, but feeble; flagellum filiform, the first three joints much lengthened, slen- der and often of an equal length; marginal cell longer than the SUOuii ass oe See etree aie hee ee ee ee (212) Orthizema Forster.
31. Clypeus medially somewhat produced and deflected at the sides.
(213) Chriodes Forster.
Clypeus not produced-medially and not deflexed at the sides.
Antenne not strongly thickened behind the middle---..............-- 32
Antenne strongly thickened behind the middle and the acuminate.
(214) Agasthenes Forster.
32. Head not much shortened, also not especially broad nor lenticular... -_ ~~~ 33
Head much shortened, very broad and almost lonticular.
First abscissa of radius usually small, scarcely longer than the transverse cubital nervure, at the most one-eighth the length of the second aDSClSSaleeree es eee ee ee Oe (215) Xenobrachys Forster.
First abscissa of radius at least one-third the length of the second abscissa.
(216) Brachycephalus Forster.
33. First abdominal segment not bent at the middle, also not elevated ___-_---- 34
First abdominal segment bent at the middle and somewhat elevated.
(217) Naétes Forster.
34. Last joint of maxillary palpi not longer than the penultimate and shorter than
NOs Meese sweeter erate ees ie ee oe (218) Ilapinastes Forster. Last joint of maxillary palpi longer than the penultimate. Scutelllumaelaterallysmarcoined only at base a-2--.2-2.ses--2 22424522 -4—- 35
Scutellum laterally margined to the apex. Spiracles of the third abdominal segment placed far from the lateral
TV AG OD Ng Me pst ar foe Seer pat o eeeeye msctae (219) Gnotus Forster. Spiracles of the third abdominal segment placed quite near the lateral TAT OUT eee ee Serer Seep rece Ss (220) Itamus Forster.
35. Metathorax with the areola toward the base, not regularly and sharply pointed, usually hexagonal or wanting, the basal area not triangular_ 36 Metathorax with the areola toward the base very regularly and sharply pointed, pentagonal, the basal area triangular.
Second and third abdominal segments sharply but finely aciculate. ; (221) Eudelus Forster. Second and third abdominal segments not transversely aciculate; wings TASC LAG ay eee eee Sears sins ot eien (222) Idemum Forster.
36 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXIII. 30 ssBasallarearand yareo lay wan Ging: eae ee eee ee (223) Phatnacra Forster.
Basal area and areola present, or at least the areola is present. Last joint of the hind tarsi hardly one-third longer than the fourth -.._ 5
Last joint of the hind tarsi hardly one-fifth longer than the fourth; anterior margin of the clypeus not truncate. Second abdominal segment with sharp well-defined thyridia. (224) Ethelurgus Forster. Second abdominal segment without thyridia-. (225) Zoophthorus Forster. 37. Transverse median nervure in hind wings broken below the middle--...----- 38 Transverse median nervure in hind wings broken at the middle. (226) Diatora Forster. 38. First three abdominal segments transversely impressed . . (227) Encrates Forster. First three abdominal segments not transversely impressed. Clypeus not separated, quite flat, anteriorly truncate or medially projecting
and feeblyomarcineds #22 -2e2 sees eee (228) Adiastola Forster. Clypeus more or less distinctly separated. Metathorax witheut two regularly formed transverse carinee - --- -- - - 39
Metathorax with two transverse carinz, but without a closed areola. (229) Isdromus Forster. 39. Second abdominal segment not finely, longitudinally aciculate-....-..------ 40 Second abdominal segment finely, longitudinally aciculate; metathorax biden- tate, the areola in the male as long as the petiolar area. (230) Ocymorus Forster. 40. Abscissa of the cubitus behind the transverse discoidal nervure so strongly bent upward that it extends parallel with the transverse cubitus. (231) Urithreptus Forster. Abscissa of the cubitus behind the transverse discoidal not parallel with the transverse cubitus. Head behindinot very much narrowed ses eee en ee 41 Head behind very much narrowed ..--------- (232) Hemiteles Gravenhorst. 41. Metathorax with the spiracular area with a sharp carina within. (233) Friplanus Forster. Metathorax with the spiracular area without a sharp carina within; first abdom- inal segment without a sharp carina extending from the spiracles to apex.
Clypeus with the anterior margin not impressed medially; abscissa of the cubitus lying behind the transverse discoidal nervure and ex- tending parallel with the second abscissa of the radius; ocelli in male not close to the eyes....------ (234) IJsochresta Forster.
Clypeus with the anterior margin medially impressed; abscissa of the cubitus lying behind the transverse discoidal nervure but strongly con- vergent with the second abscissa of the radius; ocelli in male very Close to theieyeseesss—— ae (235) Charitopes Forster.
Tribe LV. -PEZOMA CHINE
1868. Pezomachoide, Family 23, Férsrer, Verh. d. naturh. Ver. pr. Rheinl., XXV, pp. 144 and 173. 1873. Hemitelina, Tribus THomson (part), Opus. Ent., V. p. 468. 1900. Pezomachini, Tribe IV, ASHMEAD, Smith’s Insects of New Jersey, p. 569. This tribe is here restricted to species having a non-areolated meta- thorax, or at most with only a transverse apical carina. The sub- apterous females with an areolated metathorax are removed to the
NO. 1206. ICHNEUMON FLIES—ASHMEAD. on
Phygadeuonini, where, in fact, Forster had already placed some of them under different generic names, namely: Stébeutes, Pezoporus, Phyrtus, and Chamezelus. Aptesis Forster, as originally defined, seems to have included some 7f not most of these forms. The name, therefore, may have to disappear entirely, since all of the species placed here by Forster that I have had for examination belong to other genera, in a different tribe.
Agrothereutes Forster also can not be retained in this tribe. It is removed to the tribe Cryptine.
In this group the females, so faras my own observations go, seem to be always wingless or subapterous, never fully winged, while the males are most frequently fully winged, although wingless males are not rare. Both in this country and in Europe the males have been frequently mistaken and described as species of //em/teles, and at present many of them will be found in our catalogues under that genus.
The strongly inwardly curved basal nervure of the front wings, together with the broad triangular stigma, will, however, easily sepa- rate them from genuine //em/telini.
TABLE OF GENERA. Seeareneeclos: (tina Gc) mem eer eee = ae sr. nec ete oo ee Sie Pod tad Sa besa de an 5 Wingless or subapterous. Ovipositor elongate, usually longer than half the length of the first abdominal seoment, the:second, segment normal, .-.-._ 5.221.222 5-..-2.--- 2 Ovipositor much abbreviated, either scarcely exserted or so short that it does not attain half the length of the first abdominal segment.
Second abdominal segment very large, occupying most of the surface of abdomen; metathorax abruptly, obliquely truncate behind, the truncature superiorly bounded by a sharp carina; petiole very long and slender, not widened at apex... . (286) Thaumatotypus Forster.
Second abdominal segment normal; petiole widened at apex.
Metathorax sloping from the base; first joint of the flagellum longer
them phe seconder. 2-225 2585252. 2222 (237) Cremnodes Forster.
Metathorax not sloping from the base; first joint of the flagellum not longertihanitheisecond. == =. -225225... (238) Apterophygus Forster.
Deere lect Oey eect, So ey ek bao. och swe es 4 Sucre MMO Se Mins See ep Re ore OE Ae Skye th Pla eee ee 3
3. Rudimentary wings usually extending to or beyond the base of the metathorax; first abdominal segment punctured, not longitudinally aciculate, or
SUI aAbe we es ee ee © gee es, tes eee aN Yes ER (239) Aptesis Forster.
Rudimentary wings not extending to base of metathorax, often scale-like; first abdominal segment more or less longitudinally striate, longitudi-
nally wrinkled, or opaque, coriaceous -~ - - (240) Theroscopus Forster.
Wings wanting; metathorax with the apical transverse carina present; abdomen with 6 dorsal segments, the second and third large, subequal, the
first not longitudinally striate .__--- (241) Pezomachus Gravenhorst.
Pemnace Ol tne usual dengths 95) 22h ae (241) Pezomachus Gravenhorst. Race much tO MtevIAted 262 ae oe ee eo, (242) Pezolochus Forster.
5. Transverse median nervure in hind wings broken far below the middle. (240) Theroscopus Forster.
38 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXIII.
Metanotum more or less distinctly areolated, the surface irregularly nyclose. (241) Pezomachus Gravenhorst. Metathorax exareolated, with only the apical transverse carina present, the
surface coriaceous or granulate ----- (241) Pezomachus Gravenhorst. Transverse median nervure straight, not broken, the subdiscoidal nervure BOSEnES He ae se Se ER ee ee See (243) Hemimachus Ratzeburg.
Tribe V: “-HEMIGASTE RINE
This group is proposed for two genera differing from any in the preceding tribes in having a narrow, lanceolate stigma. In this char- acter it comes nearest to the two tribes which are to follow, the Cryptine and the Mesostenini, but it is at once separated from them by the areolet in the front wings being wholly absent.
The marginal cell is rather long, extending almost to the tip of the wing; the first transverse cubitus is short but distinct; the second recurrent nervure joins the cubitus beyond this vein; while the subdis- coidal nervure originates from above the middle of the discoidal nervure.
The two genera falling here may be separated as follows:
TABLE OF GENERA.
Hind wings with the transverse median nervure straight, not broken, the subdis-
coidal mervurre warm times ee eis ee See eee err 2
Hind wings with the transverse median neryure angularly broken; ovipositor short, less than half the length of the abdomen; metathorax areo-
DAOC Brix 2 eee ae te acne Lone Lee Te (244) Hemigaster Brullé.
2. Mesonotal furrows more or less distinct; metathorax incompletely areolated;
ovipositor as long as or longer than the abdomen. (245) Macrogaster Brullé.
rerbe Vas * Clover GAs
1868. Cryptoidx, Family 26, Forster, Verh. d. naturh. Ver. pr. Rheinl., X XV, pp. 144 and 186.
1873. Cryptina, Tribus (part), THomson, Opus. Ent., V, p. 468.
1894. Cryptini, Tribe IV, AsHmeEan, Proc. Ent. Soc. Wash., III, p. 278.
1900. Cryptini, Tribe V, AsHMEAD, Smith’s Insects of New Jersey, p. 570.
To this tribe belong the genuine Cryptines distinguished by the nar- row lanceolate stigma and the distinct, usually pentagonal, areolet of the front wings, although this sometimes appears quadrate from having the two transverse cubiti straight and parallel, or nearly.
In addition to the shape of the stigma, it is further distinguished from the other tribes previously defined, except the Pezomachini, by metathoracic characters. The metanotum, except sometimes with an areola, is exareolated, and is without distinct longitudinal carine, or at the most the pleural carine are alone present, the lateral longitudinal ‘arin always absent, its disk being simple, without carinz, or with one or two transverse carinee.
at ence
NO. 1206. ICHNEUMON FLIES—ASHMEAD. 39
TABLE OF GENERA.
Seen ela Re Wie ene eters racine arias oe a Saicin Salon sean se see 28
SRemeCc say mO CUCU tee nem ae Met aa tt ose we Sab cee new e seas eect 1
1. Transverse median nervure in hind wings broken distinetly below the middle,
usually far below the middle.-..--.--..-.- Sa Mate So et hee 8
Transverse median neryure in hind wings broken at, near, or somewhat. above
the middle, rarely very slightly below the middle ............- 2
2. Transverse median neryure in front wings originating either before or behind
: the basal nervure, never distinctly interstitial with it........-- 63 Transverse median neryure in front wings interstitial with the basal nervure.
MetainoraxeiOncMOniere 4 owe wees se oo aie one ks See 3
Ls SLOW eETS LE) CCC Gp es aan tgs es pee SN a el de, Mean eS 6
3. Metanotum exareolated, but with two transverse carinze, the apical carina some-
times obsolete medially, the spiracles short, oval, or rounded... _- 4
Metanotum usually with 6 more or less distinct areas above; clypeus with a slight median tooth anteriorly; disco-cubital nervure not broken by a stump of a vein; apex of seventh or eighth dorsal abdominal seg- ment with a white spot; tarsi with the fourth joint cordate, the last very large, as long as the second or longer. (246) Aritranis Forster!= Hygrocryptus Thomson. 4. Disco-cubital nervure broken by a stump of a vein. Diseo-cubital nervure not broken by a stump of a vein. Abdomen mostly red, not spotted with white. (247) Habrocryptus Thomson. Clypeus without a median tooth anteriorly but with a transverse furrow or impression; areolet large, the sides parallel..............2-2...- 5 Clypeus with a median tooth or projection anteriorly, the labrum usually pro- jecting as a ledge from beneath; pleural carinse of metathorax want- ing; dorsal abdominal segments 7-8 with a white spot. (248) Hoplocryptus Thomson. 5. Antenne normal; metathorax and sides of the thorax not striate; dorsal abdom- inal segments 7 and 8 spotted with white at apex. (249) Spilocryptus Thomson (part. ). Antenne abnormal, compressed or dilated toward apex; first joint of the fla- gellum not longer than the second; metathorax and the sides of the thorax striated; abdomen not spotted with white. (Mexico. ) (Female.) (See p. 40)....(250) Joppoceras Ashmead, new genus. (Type, Cryptus dubiosum Cresson Ashmead, manuscript. ) Metathorax usually short, sloping off from its base, or obliquely truncate, but with two transverse carinze; abdominal petiole normal, more or less broadened and bent or elbowed at apex; clypeus anteriorly truncate orslightly arcuate, but without amedian tooth; disco-cubital nervure most frequently broken by astump of a vein; abdomen red or red with black at apex, without white spots; submedian cell a little shorter than the median, rarely equal; metathoracie spiracles elonwatem eames ace sse yee Se ee ee (251) Trychosis Forster. Metathorax not short. Petiole of abdomen elongate, slender, not or only slightly thicker at apex than at base, as seen from the side straight or nearly, at most gently curved, but never distinctly bent or elbowed ...............-.- 7 Petiole of abdomen in female thickened and bent or elbowed at apex, slenderer in male; disco-cubital nervure not broken; abdominal segments 7-8 with a white spot above; metathoracie spiracles small, oval, the pleural carinse present. (249) Spilocryptus Thomson (part.).
~ co nie
us genus should be removed to the Tribe Phygadeuonini. 'This ger hould } 1 to the Tribe Phygad
40 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXIII.
7. Metathorax strongly striate, with two transverse carinze, the spiracles elongate; wings black or dark fuscous, the areolet large, with parallel sides; antennz in female broadened and compressed between the middle and apex (male) (see p. 39). -(250) Joppoceras Ashmead, new genus.
(Type, Cryptus dubiosum Cresson. ) Metathorax not striate, but with two transverse carine, the spiracles elongate; wings mostly hyaline, the areolet large pentagonal, the sides slightly convergent above; antennee filiform.
(252) Linoceras Taschenberg = Osprynchotus Kriechbaumer nec Spinola. (Type, Cryptus macrobates Grayenhorst. ) Metathorax rounded off posteriorly, with only one transverse carina—the basal ‘arina, the spiracles large, elongate; wings black, brown, or fuscous,
never hyaline, the areolet large pentagonal, with parallel sides.
(253) Joppidium Walsh.
(Type, Joppidium raficeps Walsh. ) 8. Second joint of maxillary palpi normal, never much dilated; antennze in female usually filiform, setaceous, or flagelliform.-.-.-...--...--------- 9 Second joint of maxillary palpi abnormal, much dilated, or triangular; antennze in female thickened medially, ringed with white in both sexes; meta- thorax coarsely rugose, the upper hind angles toothed or subdentate, the areola indicated but poorly defined, never distinct, the spiracles Lome sO alee e ys Bee eee age er ie oe Syke ne cine (254) Megaplectes Forster. 9. Petiole of abdomen nof cylindrical throughout, but dilated and usually bent or elbowed at apical third, where it is always more than twice wider than at base, or even thrice as wide, except in some males -.-_-- 12 Petiole of abdomen slender, cylindrical, and nearly of a uniform thickness throughout, or at the most only a little thicker at apex than at base, never twice as wide; as seen from the side it is straight or nearly, or at most slightly bent but never elbowed. Metathorax with the apical transverse carina present, the spiracles elongate; head rostriform, the malar space long. -- (255) Osprynchotus Spinola. Metathorax without a transverse carina, or at most with only a basal trans- verse carina, smooth, punctate, or transversely striate, especially posteriorly, but sometimes with a smooth, semicircular or triangular
shaped space at base, the spiracles long linear .............--.- 10 10. Disco-cubital nervure arcuate, not broken by a stump of a vein, the areolet varia-
ble*in ‘size andshapes 2 225 soe eo hae ee 11 Disco-cubital neryure broken by a stump of a vein, the areolet rather small, nar- rowed above, the transverse medium nervure originating from beyond the basal nervure; head subquadrate, not rostriform, the
malar space normal. (Africa. ) | (256) Zonocryptus Ashmead, new genus. (Cryptus sphingis Ashmead, manuscript. )
11. Metanotum without a basal transverse carina, not short.
Areolet small, triangular, the submedian cell shorter than the median; mesonotal furrows distinct, sharply defined, the middle lobe con- vexly elevated; metathorax transversely striate. (Africa. )
(257) Metarhyssa Ashmead, new genus. (Type, Metarhyssa bifasiata AShmead, manuscript. )
Areolet large, with the sides parallel, the submedian cell a little longer than the median; mesonatal furrows distinct for two-thirds the length of the mesonotum; metathorax smooth shining. (South America. )
(258) Opisoxestus Ashmead, new genus. | ( Opisoxestus ferrugineus Ashmead, manuscript. )
ont
} No. 1206. ICHNEUMON FLIES—ASHMEAD. 4]
Metanotum with a basal transverse carina, short, rounded behind, ; Areolet large, the sides convergent soméwhat above, the mesonotal furrows
rugose.
WishinGiessee= See Ae ar em eye mers (259) Distantella Saussure. 12. Metathorax with one or two transverse carine, but without a longitudinal sul- CUS etn ee i I eas eee aa, Siena oe = Si 13
Metathorax with a longitudinal sulcus or furrow, but withoul transverse carine.
; (260) Mansa Tosquinet.
13. Diseo-cubital nervure not broken by a stump of a vein; wholly without a trace
DHSUG NER EN Dern a area ert eet On ee att: bel le et 38 19
Disco-cubital nervure distinctly broken by a stump of a vein, or at least with a trace of such a vein.
Anterior tarsi in female normal, or at most with only the fourth joint cordate or emarginate, never with joints 2-4 cordate or emarginate-. ---- 14
Anterior tarsi in female with joints 2-4 short, cordate, emarginate or lobate, as well as sometimes joints in the other tarsi; cheeks, or the malar space, long; antennee filiform or tapering off toward apex (males diffi- cult to separate from Cryptus, the forehead above insertion of antennse coneave, the spiracles of metathorax larger and longer, the upper hind angles rarely dentate, while the apical transverse carina is wanting or subobsolete) ........-.------ (261) Meringopus Forster.
14. Clypeus anteriorly armed with a median tooth or projection, or angulated.. 18 Clypeus anteriorly normal, unarmed, either truncate or rounded.
Metathorax with two distinct transverse carinze, or at most with the apical carina vaguely or indistinctly defined only medially ....-.-~--- 15
Metathorax with only one complete transverse carina, or smooth without CU gene ede Se a Nees a re See ae a sare Sones osieece ves semln
15. Metathorax short, obliquely truncate posteriorly, the spiracles oval or ellip- 2 Ea (oe ee npr renee ge ery te Sere een Ls eto SUE re cae MSL 16 Metathorax not short, with the upper hind angles often toothed or with the
apical transverse carina strongly elevated laterally, the spiracles elongate, or small, short oval, or rounded.
Metathoracie spiracles large, elongate or linear, the metapleural carina indistinct or obliterated posteriorly from the basal transverse carina; median and submedian celis equal, or the latter is a little the shorter; areolet large, the sides convergent above; head transverse, narrowed behind; antennze filiform, in females most frequently ringed with white; abdominal segments 1-4 not wholly smooth, punctate or coriaceous, the spiracles of the second placed at or before the middle.
Areola of metathorax not defined; first joint of flagellum elongate, longer HAM MeESeCONGmeaa se eee en eee ee (262) Cryptus Fabricius.
(Type, Cryplus spinosus Fabricius. )
Areola of metathorax more or less defined by surrounding carine.
(263) Itamoplex Forster.
Metathoracic spiracles small rounded or short oval, the metapleural carina distinct; areolet in front wings moderately large, with the sides conyergent above; stump of vein on the disco-cubital nervure very minute or almost obliterated.
Submedian cell a little shorter than the median; metathorax with the super hind angles dentate, the spiracles small, short, oval; body marked with red, black, and white, abdomen usually with some white bandsis2.2 22.22. (264) Chromocryptus Ashmead, new genus.
(Type, Chromocryptus albopictus Ashmead, manuscript. )
Submedian and median cells equal; metathorax with the upper hind
42 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXII.
angles simple, not dentate, the spiracles small, round; thorax mostly black, marked with yellow or white, abdomen mostly red, not banded with white (see p. 39). (247) Mabrocryptus Thomson (part). 16. Submedian cell a little shorter than the median or equal to it; areolet large, with parallel sides; head transverse, not thick antero-posteriorly, the temples narrower than the width of the eyes; antennze filiform, the first three joints of the flageHum not especially elongate, the first in female a little longer than the second, in male about equal with the second; abdominal segments 1-4 wholly smooth, neither punctate nor coriaceous, the spiracles of the second placed behind
the middle; ovipositor short. (265) Idiolispa Forster = Liocryptus Thomson. 17. Metathorax finely closely punctate or shagreened, the pleural carinz distinct, the spiracles small, round; areolet moderately large, the sides con- vergent above; head transverse, the temples not well developed; antennee slender, filiform, the first three or four joints of the flagel- lum elongate, the first in female longer than the second (see p. 39). (247) Habrocryptus Thomson (part). Metathorax with (wo transverse carinze, the spiracles small, rounded, the pleu- ral carina wanting or vaguely defined posteriorly; median and sub- median cells equal or sometimes with the median cell a little the shorter; areolet with the sides convergent above; head quadrate or subquadrate, the temples full, broad; antenne filiform, ringed with white, the first joint of the flagellum a little longer than the seconds. Set Ue (266) Kaltenbachia Forster. Metathorax with only one complete transverse carina—the basal, the apical transverse carina indicated only laterally, or the upper hind angles
are toothed.
The metathoracic pleural carina complete, the spiracles oval or oblong; arcolet with sides convergent above; submedian cell slightly shorter than the median; head subquadrate; antennze not ringed with Wihite tare ee ee eee (267) Cryptoideus Ashmead, new genus.
1
ie)
(Type, Cryptus purpuripennis Cresson. )
The metathoracie pleural carina wholly absent, the upper hind angles spined, the spiracles long, elliptic; areolet large, the sides almost parallel; submedian and median cells equal; head transverse, the temples slate. ee (268) Cryptopterya Ashmead, new genus.
(Type, Cryptopteryx columbianus Ashmead, manuscript. )
19. Marginal cell rather short, the areolet large or moderate, the sides parallel or nearly, rarely small, with the sides convergent above; metathoracic spiracles small, round, short-oval, oval or ovate, never elongate or
linear; abdomen most red, black at apex, and usually spotted with
white, very rarely “wholly rede! 2722s 2 ee eee 21 Marginal cell elongate, the areolet with the sides usually convergent above, rarely parallel; metathoracic spiracles large, elongate, or linear,
never round or short oval; abdomen not spotted with white at apex. Metathorax with one or two distinct transverse carinze (sometimes wanting in males), the pleural carina more or less distinct, the upper hind angles'sometimes:dentatec: «sass eee nee eee ee 20 Metathorax without transverse carine, or at the most with the basal alone vaguely and indistinctly defined laterally, the pleural carina absent,
the spiracles very large, linear; head subquadrate, the temples rather
broad; clypeus anteriorly truncate, the labrum visible from beneath
as a semicircular ledge; first three joints of the flagellum not long,
a cn
tae ae
aes ac Set SSS
ta Pat ie
Sait Dain cate ss
tod be be pied ae. ae hte
2sde BO RE RON» Vietre.
No. 1206. ICHNEUMON FLIES—ASHMEAD. 43
not or scarcely thrice as long as thick, the first in the female not longer than the second; parapsidal furrows vaguely defined far anteriorly only; wings fuscous, maculate, or banded. (269) Compsocryptus Ashmead, new genus. (Type, Cryptus calipterus Say. ) 20. Head transverse, narrowed behind the eyes, rarely subquadrate, the malar space long; clypeus anteriorly truncate, the labrum projecting from beneath as a semicircular ledge and transversely impressed; wings usually marked with red or yellow, rarely concolorous, the median and submedian cells equal or nearly, the areolet rather large; meta- thorax with two transverse carinee, the upper hind angles toothed; abdomen coriaceous or punctate, the spiracles of the second placed a little behind the middle, those of the third much before the mid- dle; fourth joint of tarsi strongly emarginate or bilobed. (270) Callicryptus Ashmead, new genus. (Type, Cryptus fasciatus Brullé. ) 21. Clypeus normal, the anterior Inargin withow! a median tooth..........----- 22 Clypeus impressed on each side of the anterior margin and with a median tooth. (271) Cexnocryptus Thomson. Head transverse, narrowed behind or not nearly quadrate or cubical, the temiplessmeveras..widevas the eyes +2. 5. echt 2a R ae ee 23 Head cubical or subquadrate, the temples broad, full, as wide or a little wider than the eyes.
Flagellum in female usually ringed with white, the first joint not or only a little longer than the second; metathorax long, with only the apical transverse carina present, the spiracles oval or rounded; ovipositor most frequently longer than the abdomen.
bo bo
(272) Cheretymma Forster = Cratocryptus Thomson. 23. First joint of the flagellum distinctly longer than the second_............-- 24 First joint of the flagellum not longer than the second, usually a little shorter. Metathorax rather long, the basal transverse carina usually well defined but sinuate, the apical transverse carina being entirely obliterated medially; spiracles oval; areolet large, pentagonal, the sides very nearly parallel, receiving the second recurrent neryvure beyond its middle; median and submedian cells equal or the latter slightly the longer; tarsal joints strongly spinous at apex. (273) Pycnocryptus Thomson. 24. Front wings with the lower angle of the discoidal cell somewhat obtuse, the areolet most frequently large, with the sides parallel or nearly, KALE lI CONMELO ei AONE) seen nao eee. tee 2 ae aataece noe 26 Front wings with the lower angle of the discoidal cell posteriorly straight, the areolet small, the sides convergent above. Submedian vein in the hind wings not abruptly broken at the transverse median neryure but continued far beyond..................--- 25 Submedian vein inthe hind wings abruptly broken at the transverse median neryvure. Face narrowed, the cheeks short; parapsidal furrows short but distinet. (274) Hidryta Forster= Brachycryptus Thomson. 25. Metathoracic spiracles small, round, or very short oval; last joint of the hind tarsi distinctly shorter than the third or at least no longer. (275) Gambrus Forster. 26. Areolet with the sides distinctly eonvergent above........----..-...------ 27 Areolet with the sides parallel or nearly not or scarcely convergent above; malar space distinct; metathorax with the apical transverse carina some-
44 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XM.
times obliterated at the middle, the spiracles short-oval or rounded; tip of abdomen black, with one or more white spots above.
(276) Agrothereutes Forster=Spilocryptus Thomson (part).
. Metathorax not short; areolet large; tip of abdomen black, usually with one or
two white spots above.
(276) Agrothereutes Forster=Spilocryptus Thomson.
28. Metathorax with only the first transverse carina present, the second represented
by sharp ridges at the upper hind angles, the spiracles small,
rounded; first joint of flagellum elongate; apex of abdomen with a
whitespotaboOVvest-.c2s-2 foeee= eo (276) Agrothereutes Forster.
FribesVIb cMEB SOs TEI:
1900. Mesostenini, Tribe VI, Asombap, Smith’s Insects of New Jersey, p. 570.
bo ~I
This tribe is proposed for several genera heretofore placed with the —
Cryptini and which are closely allied to them. On the other hand, they present a general Avh7tus peculiarly their own, difficult to express in words, but easily recognized by the experienced eye, and which seems to me to justify their separation as a distinct minor group.
For the present, however, the only character I can give to separate them is the one used in my table of tribes, namely, the small quadrate areolet, which is sometimes nearly punctiform and often open behind, but which is never pentagonal in position, or large quadrate, as in the Cryptini. The legs, too, relatively speaking, are, as a rule, longer and slenderer, and increase more rapidly in length, antero-posteriorly, than in the others.
Ten genera have been recognized in the group, separable as fol-
lows: TABLE OF GENERA. Head with one or two spines or carinze between the antennze.-......------------ i
Head without a spine or carina between the antenne. Claws large; last joint of hind tarsi about the length of the third; basal joint of front tarsi much shorter than the tibiz; ovipositor shorter than the body 22st eee ee Thien a eee 2 Claws small; last joint of hind tarsi much shorter than the third; basal joint of front tarsi as long, or nearly, as the tibize; ovipositor usually longer than the body, or at least never shorter than the abdomen. (277) Nematopodius Grayenhorst. Thorax not, or only a little, more than twice as long as wide.......-----.--- 6
bo
Thorax three or more times longer than wide.
Mesonotum without parapsidal furrows, or at most only feebly or vaguely defined anteriorly.
Mesonotum with the parapsidal furrows distinct for at least two-thirds its length; first joint of flagellum longer than the second.__-.-.-------- 3 3. Metathorax with the upper hind angles tuberculate, dentate, or spined, the basal transverse carina alone present, distinct, the apical wanting or imcomiplete* 2a! sls else ee ere or 4 Metathorax with the upper hind angles rounded, unarmed; both transverse carine usually present, entire, although the apical transverse carina is sometimes obliterated medially, but always distinct or highly elevated laterally S32 435-5" (278) Mesostenus Gravenhorst = Stenareus Thomson.
retires 4h yest sersgere cast Sean cn warn te er ee eis
" NO. 1206. ICHNEUMON FLIES—ASHMEAD. 4
r f
o
4. Petiole of abdomen longer and slenderer, only slightly and gradually widened toward apex, not elbowed, the spiracles closer to each other than to the apex; areolet usually somewhat larger, quadrate ...........-..-._-. BS
Petiole of abdomen shorter, bent or elbowed and much widened at apical third, the spiracles usually wider from each other that to the apex; areolet smaller.
Head a little wider than the thorax; areolet closed. (279) Mesostenoideus Ashmead, new genus. (Type, Mesostenus albomaculatus Cresson. ) Head scarcely so wide as the thorax, or no wider; areolet open behind. (280) Christolia Brullé.
5. Metapleural carinz distinct to the base of the hind coxee; transverse median nervure in hind wings obtusely angularly broken by the subdiscoidal nervure near its basal third; scutellum margined at sides only STASI Oligo ot ieee mee Sis ah ae OP as Ae (281) Cryptanura Brullé
Metapleural carinze wholly wanting or at least not extending beyond the first transverse carina; transverse median nervure in hind wings straight, the subdiscoidal nervure originating from its basal fourth.
(282) Brachycoryphus Kriechbaumer.
6. Metathorax short, with only one transverse carina—the basal; hind legs much
longer than the middle and anterior pairs; areolet open behind. (283) Crypturopsis Ashmead, new genus==Crypturus Ashmead nec Graven- horst. (Type, Crypturus tecanus Ashmead. ) Head with only one spine or acute carina between the antennze; mesonotum trilobed.
sat
Metathorax with the apical transverse carina wanting or subobsolete, the upper hind angles prominently toothed*or spined; abdomen without thyridia between segments 2 and 3.......... (284) Polycyrtus Spinola.
Metathorax with the apical transverse carina distinct, prominent, the upper hind angles at most subdentate; abdomen with thyridia between seg-
TRO RS 225030 ek ae ae ee (285) Listrognathus Tischbein. Head with two spines or acute carinze between the antenne; mesonotum not rel OD eC Clyeers eee ees Ate ee tee Sate feed (286) Polyenus Cresson.
Subfamily LIT. PPVEPIUEN AD. 1859. Pimplarix Houmeren, Ofvers. Vet.-Akad. Forh., XVI, pp. 121-132. 1887. Pimplinxe, Subfamily, Cresson, Syn. Hym. North America, p. 49. 1888. Pimplariz THomson, Opus. Ent., XII, p. 1247. 1895. Pimpline, Subfamily VI, AsHmeEap, Proc. Ent. Soc. Wash., III, p. 278. 1900. Pimpline, Subfamily IIT, Asomeap, Smith’s Insects of New Jersey, p.571.
The species falling in this subfamily are readily distinguished from those previously treated of by the characters made use of in my table of subfamilies—the shape of the abdomen, the shape of the first seg- ment of the abdomen and the position of its spiracles, and the vena- tion of the front wings.
The abdomen is elongate, subcylindrical or depressed, rarely sub- compressed at apex, most frequently sessile, more rarely petiolate and always with a prominent ovipositor; the first segment is straight, rarely bent or curved, as in the /e;neumonine and the Cryptine, and with the spiracles, except in two or three cases, placed af or before
46 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXII.
the middle; while the areolet in the front wings, when present, is most —
frequently small, triangular, oblique, or rhomboidal, often petiolate, but very rarely pentagonal.
Five minor groups, or tribes, have been recognized; one being based upon Cresson’s genus Labena and its allies, and the others upon four of Forster’s so-called families—Acwnitoidw, Lissonotoide, Pimploide, and Noridoide.
These tribes may be recognized by the use of the following table:
TABLE OF TRIBES.
Head transverse, rarely subquadrate, and usually narrowed or rounded off behind, the temples not broad; mandibles always fitting close to the clypeus, not forming a kind of mouth opening.
Abdomen somewhat compressed toward apex, the ventral valye prominent, —
plowshare-shaped, or sometimes very large lanceolate; if the ventral valve is hidden, the hind coxze are abnormally long; antennz usually rather short and straight; hind legs much lengthened and usually with stout femora.
Hypopygium prominent, plowshare-shaped or lanceolate; hind coxze nor-
mal, rarely three times as long as thick..._-.------ Tribe I. Acasnrrrt. Hypopygium not prominent, hidden; hind coxze abnormally long, four or more times longer than thick. -2_- 2222222522 -22--22 Tribe IT. LABENINI.
Abdomen depressed, rarely weakly compressed toward apex, the ventral valve never prominent or plowshare-shaped; antennze longer and sometimes eroded; hind coxe never abnormally lengthened.
Abdomen smooth, without impressions and never strongly punctured, at the most alutaceous or shagreened; no lateral impressed lines on segments DE ESS 8 Se TST ae he Ra Ae ee Tribe III. Lissonorrnt. Abdomen with more or less distinct impressions and usually also strongly punctured; if smooth, alutaceous or coriaceous; always with lateral impressed lines on segments 2-5. ......----------- Tribe IV. Prcexint. Head quadrate or cubical, the temples broad, not narrowed behind; mandibles most frequently slightly projecting forward and forming, with the clypeus, a a kind of mouth opening, or the clypeus is depressed. Tribe V. Xorrp1nt.
iPribecks, ACG NEE NGL
1868. Acenitoidx, Family 17, Forster, Verh. d. naturh. Ver. pr. Rheinl., X XV, pp. 142 and 167.
1894. Acenitini, Tribe I, ASHMEAD, Proc. Ent. Soc. Wash., ILI, p. 278.
1900. Acenitini, Tribe I, ASHMEAD, Smith’s Insects of New Jersey, p. 571.
This group is distinguished from all the others by the shape of the abdomen, which is much elongated, compressed at apex, and furnished with a large, prominent, plowshare shaped, or lanceolate, hypopygium; the antenne are rather short and straight, while the hind legs are unusually long, with rather short and stout femora.
Superficially, many of the species falling in this group, especially among the males, resemble those in the tribe J/esosteninz, in the subfamily Cryptine, in colorational pattern and in having long hind
‘
“No. 1206. ICHNEUMON FLIES—ASHMEAD. 47
legs, but the shape of the abdomen and the venation of the front wings can always be depended upon to distinguish them. Sixteen genera have been recognized separable as follows:
TABLE OF GENERA.
Beaninoni chino an arecoletesa.2 sa-S2e. 280s esl - te let e le sce el eeetee]e 6 Front wings with an areolet. Areolet small, not rhomboidal; ovipositor shorter than the abdomen; if as long, then the hypopygium unusually large..-.-.-.---.-..----------. 2 Areolet large, rhomboidal; ovipositor as long as or longer than the abdomen; head as in evetartes, but readily distinguished by the longer oviposi- LOTS eee a ie etl a Oi aaa (287) Leptobates Grayenhorst. Per bodomencsessile.motlong petioliforn sees Sask hohe ce eet ens eee 3 Abdomen petiolate, or the first segment long, slender, petioliform. Areolet small, petiolet; abdomen elongate, narrowed toward base; claws pectinate yee Ps (288) Leptobatopsis Ashmead, new genus. (Type, Leptobatopsis australiensis Ashmead, manuscript. ) 3. Ovipositor not longer than the abdomen, or if longer the hind legs are very thick; antenne short and straight Ovipositor longer than the abdomen, the hypopygium unusually large, lanceo- late; mesonotum trilobed; metanotum not areolated with from two to four longitudinal carinee, the spiracles rather large, oval. (289) Coleocentrus Gravenhorst. 4. Diseo-cubital nervure without a stump of a vein; hypopygium in female large, projecting beyond the tip of the abdomen............---------- 5 Disco-cubital nervure with a stump of a vein or branch; hypopygium in female short; metanotum without carinee; ovipositor longer than the abdo- TM CMe ee tee tae See eee, ve (290) Procinetus Forster. 5. Transverse median nervure in hind wings broken at the middle; metathorax laterally coarsely rugose, the middle space punctured but shining, the spiracles large, oval; ovipositor longer than the abdomen. (291) Mesoclistus Forster. Transverse median nervure in hind wings broken far below the middle; meta- thorax short, truncate posteriorly, and completely, areolated, the spiracles small, round; scutellum and postscutellum laterally sharply margined; ovipositor somewhat shorter than the abdomen. (292) Aphanoroptrum Forster.
6. Second joint of tarsi longer than the four following joints united ...-...----- 7 Second joint of tarsi not longer than the four following joints united. ....---- 8 7. Second recurrent nervure uniting with the discoidal nervure before the very
short transverse cubital-transverse median nervure in hind wings
broken above the middle; first abdominal segment narrow, almost
three times as long as wide ---..----- (293) Crypturus Gravenhorst.
Second recurrent nervure uniting behind the transverse cubitus; abdomen a
little longer than the head and thorax united, the petiole elongate.
(294) Encardia Tosquinet.
8. Second recurrent neryure uniting behind the transverse cubitus -...--------- 9 Second recurrent nervure uniting before the transverse cubitus.
Anterior and middle claws cleft before the middle, the hind claws simple, the middle and hind tibice with 2 apical spurs; transverse median nervure in hind wings broken at or a little above the middle; hind legs much lengthened; ovipositor as long as the body.
(295) Arotes Gravenhorst.
48 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXIII.
9. Mesonotum without deep parapsidal furrows .---..--.----=-----.+.-- pone Tf Mesonotum with deep parapsidal furrows. Disco-cubitalineryurevuith a stump) of a, vein S22 sn sae ee ee 10 Disco-cubital neryure without a stamp of a vein. (296) Phenolobus Forster. 10. Second abdominal segment longer than wide at apex, the hypopygium in female
small and placed far away from tip of abdomen; coxze much length-—
ened; ovipositor somewhat prominent -- - - - (297) Collyria Schiédte.
Second abdominal segment not longer than wide at apex; hypopygium in female very large and projecting beyond tip of abdomen.
(298) Chorischizus Forster.
11. Metathorax elongate, quadrate, rugose, with a vaguely defined areola; head
large, much longer than wide, slightly narrowed back of the eyes,
the malar’spacelongse= === shee anne (299) Hieroceryx Tosquinet. Metathorax and head quite differently shaped.
Metanotum with a well-defined transverse carina; hind femora much thick-
ened; recurrent nervures very close. Femora not compressed, the tibize normal; claws of four anterior feet with a blunt tooth before tip ..........--- (300) Acenites Latreille. Femora compressed, the tibiee bent, narrowed at base, the tarsi long, the posterior dilated, the last joint not merassated, claws simple. (301) Acronus Tosquinet. Metanotum without a transverse carina; hind femora not especially thick- ened; recurrent nervures widely separated. (302) Asthenomeris Forster.
Tribe ihe “LE ABE hers
1900. Labenini, Tribe Il, AsomMeab, Smith’s Insects of New Jersey, p. 571.
This group is proposed for three genera found only in the American fauna. In the shape of the abdomen it bears a superficial resemblance to some of the Acawnitini, but the hypopygium is hidden, neither prominent, plowshare shaped nor lanceolate, while from them and the other tribes it is readily separated by the abnormally long hind coxe, which are four or more times longer than thick.
The characters made use of in the following table may be depended upon to distinguish the genera:
TABLE OF GENERA.
Cheeks armed with a strong tooth behind 2 Cheeks normal, unarmed.
Face rugoso-punctate; metathorax completely areolated; areolet in front wings large, pentagonal, the third discoidal cell shorter than the second; trans- verse median neryure in the hind wings broken slightly above the middle; first abdominal segment not especially long, very gradually widened toward apex, the spiracles placed before the middle_......-- (303) Labena Cresson.
Face finely coriaceous; metathorax long, exareolated, but with a transverse carina before the apex; areolet pentagonal, the third discoidal cell much longer than the second; transverse median neryure in hind wings broken below the middle; first abdominal segment very long, the spiracles placed a little behind themiddle. 23-52. 8 Oe (3804) Nonnus Cresson.
2. Face smooth imponctate; metathorax areolate at base only, the areola wanting;
) ; {
ee ere hee
No. 1206. ICHNEUMON FLIES—ASHMEAD. 49
areolet large, pentagonal, the third discoidal cell longer than the second; transverse median nervure in hind wings broken at the middle; first abdominal segment very long and slender, not at all or only slightly widened at apex, the spiracles at or very near the middle... __- (305) G'rotea Cresson.
iribetl ly siSsONOTIN I.
1868. Lissonotoidx, Family 16, Forster, Verh. d. naturh. Ver. pr. Rheinl., XXV, pp. 142 and 166.
1894. Lissonotini, Tribe II, Asumean, Proc. Ent. Soc. Wash., III, p. 278.
1900. Lissonotini, Tribe III, Asamean, Smith’s Insects of New Jersey, p. 371.
This tribe is distinguished from the two aforementioned by the shape of the abdomen, which is depressed, nof compressed at apex, and without a prominent hypopygium; by the longer antenne; and by coxal and venational characters.
It approaches nearest to the tribe P/mplin7, but in that tribe the abdomen, although sometimes smooth, is usually strongly punctured, and has always more or less distinct transverse or oblique impres- sions, or lateral impressed lines on segments 2-5. In the Lissonotini, on the contrary, the abdomen is smooth, or at the most alutaceous or shagreened, but never strongly punctate, always w/thout transverse or oblique impressions, and never with lateral impressed lines on seq- ments 2-5,
The 31 genera falling in this group may be separated as follows:
TABLE OF GENERA.
Metathorax not areolated, at the most with a transverse apical carina, or with two
Median loneHUCInAl CARING. ...s2.-: 1 Jess oh. bes. ees eens 3 Metathorax more or less indistinctly areolated. BromtiavnosiiuuaAneareoleterewne jos. sods JUN eh he 2
Front wings without an areolet.
Abdomen sessile, with the first three segments rugulose, the ovipositor short; clypeus large, swollen at base; flagellar joints of male antenneze normal, not eroded; eyes hairy.
(806) Hybophanes Forster= Cidemopsis Tschek.
2. Areolet oblique, elongate, subtrapezoidal-.-- (807) Pseudaccenites Kriechbaumer.
Heaamen sessile, never petiolate ...:........2s.--4-+--2s-s0-e-e- e-- eee ee 4
Abdomen petiolate, head transverse; metathorax exareolate, without a transverse apical carina.
Abdomen elongate, subfusiform, as seen from the side toward apex, clavate; marginal cell in front wings large, oblong; the areolet small, sub- triangular and subpetiolate; legs elongate, slender.
(808) Atropha Kriechbaumer.
Abdomen toward base flat, rugose, the first segment flask-shaped, distinctly and longly petiolate; marginal cell in front wings not large, the areo- let longly petiolate, the outer nervure faint or subobsolete; legs nor- mal; antenne not tapering toward apex, the last joint nearly as long as the three preceding joints united; ovipositor nearly as long as the abdomen.
(309) Taschenbergia Schmiedeknecht.
Proc. N. M. vol. xxiii—-4.
i }
50 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXII.
4. Areolet always present, rarely incomplete, that is, open behind. ---.-.------ 6 Areolet wholly wanting. Metanotum with two distinct longitudinal carinze in both sexes.-------- 5) Metanotum without longitudinal carinz. Claws closely, longly pectinate; Man Giles | Did enitales a= sesso ee aaa ee (810) Asphragis Forster. . Clypeus normal; abdomen rather smooth, the first segment flat, longer than wide; ovipositor almost as long as the body; claws simple, not toothed; eyes bare; third joint of the flagellum at tip and the fourth joint at base in male eroded. (311) Lampronota Haliday=Cylloceria Schiddte. 6: Body not..especially, hairy ac .cieenbs so seeece - 22 beatiese> ane eee 7 Body, and especially the head, covered with shaggy gray or black hairs.
Metathorax coarsely punctured; eyes widely separated; abdomen in female somewhat compressed at apex, the first segment somewhat longer than wide, rugulose, the following smooth; claws simple, the onychia very amall-.'20= 22 2ess-2 2. ae aoe (312) Arenetra Holmgren.
7. Metathorax with a distinct transverse apical carina, or at least distinctlaterally. 11 Metathorax without a transverse apical carina. Claws simple, neither toothed nor pectinate...........---------i------ 9 Claws thickly and usually longly pectinate, never simple.--..---------- 8 8. Antenne long and slender, the last joint twice as long, or nearly, as the preced- ing; abdomen smooth, the first segment without carinee, the spira- cles of the second placed close to the lateral margin.
Transverse median nervure in hind wings broken very far below the middle; eyes not quite extending to the base of the mandibles, the malar space being fully as long as the pedicel, the latter being obliquely truncate from beneath; submedian and median cells in front wings
; COU al Rae eer cc eeton ele mee cere (313) Phytodietus Gravenhorst.
Transverse median nervure in hind wings broken at or a little above the middle; eyes extending to base of mandibles without a malar space; submedian cell in front wings distinctly shorter than the median. (New Zealand.) .........-.- (314) Huctenopus Ashmead, new genus.
(Type, Euctenopus zealandicus Ashmead, manuscript. 9. Metapleural carinz: present, strong and long. .< 4. -.. 2. -4 4 eee 10 Metapleural carinze wanting.
Abdomen subpetiolate, smooth, shining, and gradually narrowed toward base; antennze long and slender, tapering toward apex, the terminal joint in male shorter than the penultimate; submedian cell not longer than the median s..-2:<-----5- se (3815) Aphanodon Forster.
Or
Abdomen distinctly sess.le, the first segment aciculate, segments 2-5 quad- —
rate, the following wider than long, all finely punctate at base, but smooth and shining at apex; antennz elongate, slender, setiform, but shorter than the body ‘2-222. -22.22--2 (316) Nadia Tosquinet. 10. Abdomen sessile, depressed, finely coriaceous; areolet in front wings pentag- onal, the submedian cell longer than the median, the disco-cubital nervure not broken by a stump of a vein. Male. (317) Trevoria Ashmead,! new genus. (Type, Trevoria yukatatensis Ashmead, manuscript.) 11. Claws simple; mot pectinate: 222 s82 ls es Sus See eee ee ee eee 21 Claws pectinate. Claws shorter, not thickly pectinate, usually briefly pectinate toward
base.!2. 22h ecu ue eeetet ale ascd- eee eee en ane ee meee 12
Claws long, strong, and usually but not always thickly pectinate....--- 17
12. Flagellum entirely composed of cylindrical, closely united, almost inseparable JOIMts:tOvipoOsitonilong ee so-so ee eee ee 15
1 Jn honor of Prof. Trevor Kincaid.
A OEE BOL MEL DEE
{
No. 1206. ICHNEUMON FLIES—ASHMEAD. Si
Flagellum with the joints composing uke apical half distinctly ne ovi- positor at the most as long as the abdomen.
Last half of the flagellum in the female with knob-like joints, appearing quite different from the basal half; the knob-like apical joints are as wide as long, almost rhomboidal, seen from beneath angulate, the last joint narrower and scarcely as long as the preceding.
(818) Xenacis Forster.
The apical third only of the flagellum with distinctly separable joints; the terminal joints are also not knob-like, but only faintly compressed, above and beneath rounded, the last joint wider and as long as the two preceding joints united; abdomen perceptibly narrowed toward
base, subpetiolate._..........---- (319) Cruptopimpla Taschenberg. 3. Frons without horns... ----- RE aes ee eee See ata Cee oases Se Seine 14
Frons with two horns.
Areolet longly petiolate; hind wings with the transverse median nervure broken below the middle; mesonotum without trace of furrows; scu- tellum laterally not margined; metapleural carina distinct posteriorly, the spiracles long, linear; abdomen with the first segment laterally toward base with two strong carine, the spiracles placed before the middle and distinctly visible from above --.-(320) Diceratops Forster.
14. Clypeus not impressed; areolet usually petiolate ....................-.---- 15 Clypeus at base posteriorly strongly impressed, the impression so covered with long hairs as to form a tuft; metanotum in female with a weak,
transverse apical carina, stronger in male; metapleural carina faint,
nearly obliterated; areolet sessile, irregularly pentagonal; spiracles
of second abdominal segment placed close to the base; claws
SUIT pee ae aes Mek ees Wye le leh ou: (321) Ensimus Forster.
15. Metapleura not separated from the metanotum by a carina, the spiracles large, elongate or linear; areolet longly petiolate; transverse median nerv-
ure in hind wings broken below the middle; face more or less
swollen; mesonotum without trace of furrows; scutellum laterally
margined only at base; first abdominal segment smooth, the spira-
cles placed before the middle; claws with abbreviated teeth and
loneapmaticss:s2tose 826 ec esos toluene (322) Zyzeuctus Forster.
Metapleura separated from the metanotum by a carina, the spiracles round or SHOT te Oyss Eee mee ere MES a ter Hie of erase, SOR AEN ORES EO eee 16
16. Claws distinctly but not closely pectinate--.--.----- (323) Lissonota Gravenhorst. 17. Areolet petiolate, rarely sessile; disco-cubital nervure angulate or angularly broken and usually with a stump of a vein -.-.....----..----- 18
Areolet sessile; disco-cubital nervure bowed or strongly curved, never angulate and without a stump of a vein. Claws shortly pectinate; transverse median neryure in hind wings broken
below the middle- - ---- (824) Meniscus Schiddte=Amersibia Forster. 18. Metathorax normal without longitudinal CANN erie see See oe han eet ce 19 Metathorax with 6 longitudinal carine -...-.------ (325) Phidias Vollenhoven.
19. Frons above the antennze normal, not at all impressed and without peculiar fovese; metapleural carinzee wanting or only faintly indicated at LOU me eet Sens aE EUIM gos SoS eon 20
Frons above the antennze impressed or concave, with the margins swollen on each side; metapleural carinz distinct; transverse median nervure
in the hind wings angularly broken below the middle; claws stout,
pectinate, but not thickly; first abdominal segment with two carinz
at the basal third... (326) Bathycetes Forster=Bathynophrys Forster.
20. Areolet sessile; metathorax with the apical transverse carina present, distinct. (Hemialel ie. eee Ne oe eek ee (327) Alloplasta Forster.
52
23.
24.
25.
PROCEEDINGS OF THE NATIONAL MUSEUM. VOL, XXIII.
Areolet petiolate. Areolet small, oblique, open behind; metathoracic spiracles small, oval; claws strongly pectinate--......------ (328) Ctenopimpla Cameron.
- Ageolet petiolate-or subpetiolate o<ci.2 33. fast Dae ee eee eee 24
Areolet sessile. Areoletitriangular or rhomboidal 2. 2-.-2 232: jee eb eee eee 22 Areolet irregularly pentagonal. Clypeus at base posteriorly strongly impressed, the impression often so covered with long hairs as to form a tuft --.(321) Ensimus Forster.
2. Disco-cubital nervure strongly curved, but not broken by a stump of a vein. 23
Disco-cubital nervure angularly broken a little before the middle.
Transverse median nervure in hind wings broken a little below the middle; clypeus anteriorly rounded; claws very long, not pectinate, but ciliate with bristles within. (Male) (p.51) (327) Alloplasta Forster.
Median and submedian cells of an equal length; transverse median nervure in hind wings obtusely angularly broken near the apical third. (South America) CS222225 ee (329) Epimecoideus Ashmead, new genus. (Type Epimecoideus apicalis Ashmead, manuscript. ) Median and submedian cells unequal; transverse median neryvure in the hind wings broken below the middle. . Clypeus prominent, separated and in outline semicircular; metathoracic spiracles short oval; areolet oblique, rhomboidal. (330) Pimplopterus Ashmead, new genus. (Type Pimplopterus alaskensis Ashmead, manuscript. ) Disco-cubital nervure strongly curved or bent, but never angularly broken, and without a stamp lof a vein. 2: cee etsatts Geet es ee eee 25 Disco-cubital nervure angularly broken, or with a stump of a vein.
Disco-cubital nervure angularly broken at the basal third.
(331) Stenolabis Kriechbaumer.
Disco-cubital nervure not angularly broken but still broken by a stump of a vein near its middle; metathoracic spiracles elliptical, thrice as long as wide; abdominal segments all longer than wide.
(332) Meyva Cameron. Metanotum with two delicate parallel, or nearly, carinee down the center, the ; spiracles rounded; abdominal segments 1-3 longer than wide, the first more than twice longer than wide, 5-6 wider than long, the last very short; transverse median nervure in hind wings broken below the middle -...--- (333) Harrimaniella Ashmead, new genus. (Type, Harrimaniella yukakensis Ashmead, manuscript.) Metanotum without such carinee, the spiracles small, rounded; third abdominal segment a little wider than long, the first narrowed toward base more than twice longer than wide at apex. (See p. 51.) (319) Cryptopimpla Taschenberg.
Tribe IV... PIMPGINE
1868.. Pimploidx, Family 15, Forster, Verh. d. naturh. Ver. pr. Rheinl., X XV, pp. 142 and 162.
1894. Pimplini, Tribe III, AsHmEap, Proc. Ent. Soc. Wash., III, p. 278.
1900. Pimplini, Tribe IV, AsHmrap, Smith’s Insects of New Jersey, p. 572.
In this tribe are to be found some of the largest, if not the largest species, of all the parasitica. The group is a most extensive one, and includes several well-known
Ra woe Seti By ren wee A Dinlaeese? oy moe
No. 1206. ICHNEUMON FLIES
ASHMEAD. 53
genera of a world-wide distribution, and among which are such con- spicuous genera as Léhyssa and Thalessa, commonly known as the Long-Stings. The species belonging to these two genera destroy the xylophagous saw-flies belonging to the genera Sirex, Paururus Tremex, ete., which - bore and live in the interior of various forest trees, and in order to reach their hosts their ovipositor has become enormously developed, attaining sometimes the length of 5, 6, or more inches. Undoubtedly this group is closely allied to the Z2ssonotin7, andsome authorities would unite the two groups; but I agree with Forster in thinking the P’m- plini distinct since they may be readily separated by the sculpture of the abdomen, the transverse impressions or furrows on the segments and particularly by the lateral impressed lines on segments 2 to 5, which are never present in the Lssonotiné. Sixty-one genera have been recognized in the group separable as
follows: TABLE OF GENERA. Sanath 700 LADS VELSeLy: TUSOSE = -.2- == -s~ = i <-- 23 oo anne eee eecenseeceesus 4 Mesonotum transversely rugose. ENCCOLCUMMEOMER WINS LCSCN UG o.oo cn.n eens tate we ences Se acts cao sececchesee 2
Areolet in front wings wanting. (334) Epirhyssa Cresson= Rhyssonota Kriechbaumer. (Type, Epirhyssa speciosa Cresson. ) 2. Metathorax not areolated; head without a carina between the antennze ___...- 3 Metathorax areolated; head sometimes with a carina between the antenn.
Head with a strong carina between the antennze; transverse median neryure straight, not broken, the subdiscoidal nervure originating from the median vein far beyond the apex of the submedian cell.
(335) Apechneura Kriechbaumer. (Type, Rhyssa terminalis Brullé.) Head without a carina between the antennze. (336) Certonotus Kriechbaumer. (Type, Certonotus varius Kriechbaumer. ) 3. Transverse median neryure in hind wings broken far above the middle, or the subdiscoidal neryvure originates from the angle formed by the trans- verse median and the median veins.
Head with the temples broad, well developed, nearly the width of the eyes; hind tarsi much elongated, very much longer than their tibize; abdo- men in female compressed toward apex.
Clypeus anteriorly medially lengthened or unidentate; abdominal seg-
ments at apex either rounded or emarginate.
(337) Rhyssa Gravenhorst.
(Type, Ichneumon persuasorius Linneeus. )
Clypeus anteriorly truncate; abdominal segments in male smooth, with segments 3-7 at apex emarginate or deeply excayated.
(338) Thalessa Holmgren.
(Type, Ichneumon clavator Fabricius. )
Head with the temples very narrow, scarcely half the width of the eye; hind tarsi normal, hardly longer than their tibize; abdomen depressed.
(339) Lylarmes Cameron. (Type, Lytarmes maculipennis Cameron.)
|
54
PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXIII.
Second abdominal segment in female usually much longer than wide, rarely quad- rate at apex; Ovipositor most frequently longer than the body... 5
Second abdominal segment in female transverse or quadrate, seldom a little longer than wide; ovipositor usually shorter than the body ----- 9
Antenne normal, the joints 3 to 5 outwardly not serrate.............-.----- 6
Antenne with the joints 3 to 5 outwardly serrate.
Abdomen with segments 1-7 longer than wide, with indistinct lateral swell- ings; transverse median nervure in hind wings faintly broken below the middle; thorax for the greater part red.
(340) Troctocerus Woldstedt.
(Type, Troctocerus elegans Woldstedt. )
Abdomen in both sexes nearly of an equal width throughout, the sides parallel or
nearly, the sculpture of the anterior segments not different from
that of the posterior segments, or only slightly; disco-cubital nery-
ure.straight, curved or “angulated... <2: 35-5 .-p2 545. eee 7
Abdomen in female spindle-shaped or tapering toward base and apex, the sides not nearly parallel; in male straight, broad cylindrical.
Segments 2-3 in female or 2-5 in male with oblique furrows; disco-cubital neryure angulate and with a short stump of a vein; ovipositor some- what shorter than the body-.--- (341) Atractogaster Kriechbaumer.
(Type, Atractogaster semisculptus Kriechbaumer. )
7. Claws in female simple, without a tooth at base; last joint of hind tarsi at least
8.
o
thrice as long as the preceding; male with the inner margin of the eyesidéeply emarpinate 9" sao oe eee ee 8 Claws in female cleft or with a strong tooth at base; last joint of hind tarsi longer than the preceding; male with the inner margin of the eyes not distinctly emarginate. Transverse median nervure in hind wings broken above the middle; metatho- racic spiracles oval or elongate--...---- (342) Ephialtes Gravenhorst. (Type, Ephialtes tuberculatus Gravenhorst. ) Transverse median nervure in hind wings broken at or below the middle; meta- thoracic spiracles small, round. (343) Calliephialtes Ashmead, new genus. (Type, Pimpla xanthothorax Ashmead.) Metathorax smooth, shining, without punctures; areolet in front wings rhom- boidal, not petiolate; transverse median nervure in hind wings broken above the middle .........-.---- (344) Perithous Holmgren. (Type, Pimpla divinator Gravenhorst. ) Metathorax punctate, and medially irregularly, transversely rugulose, evanescent toward the sides; a subsemicircular area posteriorly; areolet in front wings small, petiolate; disco-cubital nervure arcuate. (345) Opisorhyssa Kriechbaumer. (Type, Opisorhyssa flavopicta Kriechbaumer. ) Abdomen distinctly punctate, or coriaceous, or at least never perfectly smooth,
shining, or impunctate ss. stesso ee ee eee ee 15 Abdomen perfectly smooth, shining, impunctate, or at the most feebly aluta- ceous. Areolet always present:-......2..--32---22-25- de ete Mays Ne RS So ceeenee slit Areolet: Wanting = 21s 2-jo. Sink 25. Be Se ee eee ee 10
10. Eyes very large, occupying the whole sides of the head, the temples usually flat;
ocelli large, prominent; claws"with a tooth toward base beneath. Prothorax narrowed into a neck anteriorly; eyes convergent anteriorly; temples iat onOpliques=.s---5-—— == === as (846) Epimeces Brullé. (Type, Epimeces bicolor Brullé. )
ee a SO Ce Ne en eee ee ard eee) ee ie Sel) Os
So cma sll BOR Cente te le
No. 1206. ICHNEUMON FLIES—ASHMEAD. 55
Prothorax not narrowed into a neck anteriorly; eyes not convergent anteri- orly; temples neither flat nor oblique--_-- - (347) Hugalta Cameron. (Type, Hugalta strigosa Cameron. ) 11. Eyes with the inner margin emarginate or subemarginate; middle vein in hind ViMLayeRS} lis HU AYES Ko) HaYe) lorVst Re es ae ee ee 5 erg ae 12 Eyes with the inner margin entire, not at all emarginate; middle vein in hind wings sometimes obsolete at base. Middle vein in hind wings distinct to base; metathorax areolated. Metathorax irregularly transeversely striate, and with a shallow median furrow on the basal three-fourths. (348) Pseudeugalta Ashmead, new genus. (Type, Eugalta spinosa Cameron. ) Metathorax smooth and polished, without a median furrow, the spira- cles small, round; disco-cubital nervure not broken by astump of a : vein; transverse median neryure in hind wings broken below the TMI CIE Hee eee oe (Africa) (349) Zonopimpla Ashmead, new genus. (Type, Zonopimpla albicincta Ashmead, manuscript. ) Middle vein in hind wings toward base obselete; metathorax more or less distinctly areolated, the areola and the petiolar area confluent; hind femora not much swollen; ovipositor longer than the abdomen. (350) Idiogamma Forster. (Type, Idiogamma euryops Forster.) 12. Mesonotum with sharply defined parapsidal furrows, which converge and meet before attaining the base of the scutellum; claws stout, simple; metathorax with the upper hind angles toothed, without an areola, but with a distinct petiolar area; hind femora with a small tooth beneath toward apex; transverse median neryure in hind wings broken far above the middle.-.. (351) Lissopimpla Kriechbaumer. (Type, Lissopimple 8-guttata Kriechbaumer. ) Mesonotum without distinct parapsidal furrows, either entirely wanting or only vaguely defined anteriorly; hind femora more or less thickened; ovipositor at most never larger than the abdomen, usually shorter. Clangarcimaplewmnotspechinatetes sans sce saeco ae eee ae ee ome cen = oe 14 Mlaweverylarce, strongly pectinate : . = .2.------.- se. eese5------ 22 13 13. Metathorax with a distinct areola and a petiolar area.. (352) Theronia Holmgren. (Type, Pimpla flavicans Fabricius. ) Metathorax without either an areola or a petiolar area. Head normal, the malar space short -...------ (353) Neotheronia Krieger. (Type, Theronia tolteca Cresson. ) Head subrostriform, the malar space long. (See p. 57.) (3866) Echthromorpha Holmgren. 14. Metathorax exareolated but with a strong transverse apical area, the upper hind angles dentate or tuberculate; hind femora unarmed. (New JASON En 0X0 C9) eee ae ea (354) Allotheronia Ashmead, new genus. (Type, Allotheronia 12-guttata Ashmead, manuscript. ) 15. Last joint of antennee not longer than the two preceding joints united; last joint of hind tarsi two or more times longer than the preceding joint. 16 Last joint of antennz large, oblong, longer than the two preceding joints united; last joint of hind tarsi not fully twice as long as the preceding joint. Face clothed with long silvery hairs; scutellum only slightly margined at sides; metathorax areolated..-.........------ (355) Stilbops Forster. (Type, Pimpla veluta Gravenhorst. ) 16. Clypeus distinctly separated; eyes rarely hairy -.......-.----------------- Clypeus not separated; eyes either hairy or bare, with their inner margin entire, or atimostoniy shehtly;emaroimate. 52-2. --..-55-.2---------- 17
56 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXIII.
17. Eyes hairy; mesonotum with distinct parapsidal furrows; abdomen narrow, the first segment bicarinate, the last ventral segment short; ovipositor not long; wings without an areolet ... (356) Schizopyga Gravenhorst.
(Type, Schizopyga podagrica Gravenhorst. ) Eyes bare; mesonotum with the parapsidal furrows indicated only anteriorly; abdomen as in Pimpla, the first segment bicarinate. Metathorax not areolated, the spiracles long; ovipositor longer than the abdomen; claws with a strong angular tooth at base. > (357) Hemipimpla Saussure. Metathorax areolated, the spiracles small, oval or elliptic; ovipositor shorter than the abdomen; claws simple withowt a tooth at base; areolet in front wings wanting (Africa). } (358) Neopimpla Ashmead, new genus (Type, Neopimpla abbottii Ashmead, manuscript. )
18. Wingawiih an areolst ic. 2. Sos oe Se Se ee oe eee 19 Wines without amiareoletsice SSeS ae ase eee eee een oc sean 31 19. Claws not pectinate, or only faintly and indistinctly. ........-.--..-.------ 20
Claws strongly pectinate.
Metathorax with a transverse apical carina, the spiracles linear; first abdominal segment bicarinate; stigma narrow, the radius originat- ing before its middle, the areolet large, tetragonal, briefly petiolate; clypeus conyex; mesonotum with furrows anteriorly.
(359) Odinophora Forster. 20. Abdominal segments with transverse impressions, especially laterally near apex onsegmients: 2-4 ei ines OER hee ee Ce eo eee 21
Abdominal segments with strong oblique impressions or grooved lines.
Areolet large, tetragonal; abdominal segments 2-3 only with oblique impressions; transverse median neryure in hind wings broken at tila ara Cl Gh ee tr Pe ee te ee (360) Dyspetes Forster,
Arrolet small, petiolate, not rhomboidal; abdominal segments 2-4, with oblique impressions.
Forehead with two horns; seutellum black; abdomen banded with Wihtte.c seas foe ones ee eRe (361) Hoplitophrys Forster. (Type, Glypta brischkei Holmgren. )
Forehead with one horn; scutellum and the extreme apical margins of the segments yellow; transverse median neryure in hind wings
broken above the middle -.--.....-..------ (3862) Teleutea Forster.
(Type, Glypta striata Gravenhorst. )
21: Metathorax not areolated: Yosh Se aan eae ae ee ee eee enee eer ne aes 22 Metathorax areolated, or at least with a complete areola and a petiolar area.
Scutellumimormalse2e 22. oe ee (363) Delomerista Forster.
(Type, Pimpla mandibularis Grayenhorst. )
Seutellum conically elevated and margined at sides; transverse median neryure in hind wings broken far above the middle.
(364) Xanthopimpla Saussure.
(Type, Xanthopimpla nova Saussure. )
Clypeus not separated from the face at base.
Clypeus anteriorly semicircularly emarginate; abdominal segments 2-5, with deep, transverse furrows at base and apex, which are united with an impression along the sides; metathorax smooth, exareolated, without a trace of carinse; claws strong, with a large tooth or lobe at base; transverse median neryure in hind wings broken very far
Mee hots
ICHNEUMON FLIES—ASHMEAD. Da
below the middle; areolet in front wings oblique, rhomboidal.
(Lower Siam) ......-... (865) Erythropimpla Ashmead, new genus (Type, Erythropimpla abbottii Ashmead, manuscript.) 23. Clypeus impressed anteriorly at apex ....-.-...-- ate A ee oss clone Dy
Clypeus not impressed anteriorly at apex. First abdominal segment with a hump-like elevation toward the apex.. 24 First abdominal segment normal, without a hump-like elevation.
Head subrostriform, with a broad malar space; eyes slightly convergent anteriorly and subemarginate within; areolet petiolate, the sub- median cell longer than the median; transverse median neryure in hind wings not broken, the subdiscoidal neryure interstitial.
(366) Hchthromorpha Holmgren. (Type, Echthropimpla maculipennis Holmgren. )
Head normal, not subrostriform; eyes entire, not convergent anteriorly; transverse median nervure in hind wings broken, the subdiscoidal neryure not being interstitial .........-.-- (3867) Tromatobia Forster.
(Type, Pimpla variabilis Holmgren. ) 24. Head normal; abdominal segments 2-4, with a transverse impression laterally near apex and with oblique lateral impressions at base; submedian cell longer than the median, the disco-cubital nervure not broken, the areolet rather large, sessile; metathoracic spiracles oval. (Gilaiwals) aes sees (368) Glyptogastra Ashmead, new genus. ) (Type, Glyptogastra hawaiiensis Ashmead, manuscript. ) 25. Transverse median nervure in hind wings broken, the subdiscoidal nervure OUP RS bial eames eee eee Se ee uer eee BONE & boas ase 26 Transverse median nervure in hind wings straight, not broken. Metathoracic spiracles small, round; hind femora normal; second absciss¢ of the radius straight, the median and submedian cells equal in
lenpibyemer er oms iS oo ets sete oe ISS: (869) Tromera Forster.
(Type, Pimpla pomorum Ratzeburg. )
MECMNEAC GHPILACLeSMrOUNG =. 2 i528 2 2 Soe esto lok eessle fees ee. 29 Metathoracie spiracles linear, oval or reniform.
Clawsunitemalenzthya, tooth beneath -....-..--..---.-------------:-- 28
Saweindemale simple, without a tooth ........-.00.5-----4--2224-4--- 27
27. Metanotum with two elongate more or less distinct areas; head subrostrate;
antenne with the joints toward apex nodosely incrassated. (Male. )
(366) Echthromorpha Holmgren= Polyamma Kriechbaumer.
Metanotam without areas and without a transverse apical carina; head and
antenne normal; transverse median nervure in hind wings broken
abovesmemmddless. 52 22c2s0c-e5e+52-5- (370) Pimpla Fabricius.
Metanotum exareolated, but with a distinct transverse apical carina; head
and antennz normal; areolet trapezoidal, subpetiolate; transverse
median nervure in hind wings obtusely angularly broken above
tocemnnid dle meres =e nee ety fil (371) Notopimpla Kriechbaumer.
(Type, Pimpla terminalis Brullé. )
28. Eyes in both sexes deeply emarginate within; lateral ridges of the mesonotum
extending on to the scutellum; ovipositor directed upward at tip.
(372) Apechthis Forster.
(Type, Pimpla rubata Grayenhorst. )
Eyes not, or scarcely, emarginate within; lateral ridges of the mesonotum nol extending on to the scutellum; ovipositor at tip straight.
(373) Everistes Forster.
(Type, Pimpla roborator Grayenhorst. )
J
58 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXIII. 29 Claws nvihoutatoothibeneath 222 Jee ese) oe eee eee eee 30,
30.
31.
33.
34.
30.
. Front femora not especially thickened, not excised........---..----------- 33m
Claws with a tooth beneath at base.
Transverse median nervure in hind wings broken far above the middle and almostiat:a right angle 24) 22 See (374) Iseropus Forster. (Type, Pimpla holmgreni Schmideknecht. ) Transverse median nervure in hind wings broken at or below the middle, seldom a little above, but usually at a very obtuse angle. &
(375) Epiurus Forster.
(Type, Pimpla brevicornis Gravenhorst. ) |
The sharp lateral ridges of the mesonotum extend on to the sc utellum; trans- verse median nervure in hind wings broken before the middle, but | always at amight angle. 3.2.2 2222 52. 222 (376) Itoplectis Forster. (Type, Pimpla maculata Gravenhorst.) The sharp lateral ridges of the mesonotum do not extend on to the scutellum; transverse median neryure in hind wings broken at an obtuse anglall atiomberore the umiddlesmissssss=s se ao (377) Eremochila Forster. (Type, Pimpla ruficollis Gravenhorst.) — Transverse cubital nervure variable, rarely much longer than the basal abscissa on theicubitus» scutellumyroundedme ss 455 eee. eee 32 Transyerse cubital nervure much longer than the basal abscissa of the cubitus, — i.e., the part lying between the disco-cubital nervure, or first recur- rent, and the second recurrent; scutellum tetragonal, truncate pos- teriorly, marked with yellow; segments 2-4, with two oblique lines and with a transverse line before the apex; ovipositor shorter than thesabdomens 23: Sea eee (378) Lycorina Holmgren. | (Type, Lucorina triangulifer Holmgren.) —
Front femora gradually swollen before the middle to the tip and excised, their | tibize bent at the base, the last joint of tarsi stout with strong claws; — abdomen narrow, smooth, the first segment longer than wide, bicarinate; ovipositor scarcely as long as the first segment.
(379) Colpomeria Holmgren. (Type, Colpomeria levigator Holmgren. )
Abdominal segments 2-4, without oblique impressions; claws simple or rarely — pectinate. cues 38. 2b Se RL Se eee 36
Abdominal segments 2-4, with oblique impressions or grooved lines; claws pecti- nate, rarely simple.
oe oe RLS Dee ee ee asniteh pebenbinthien hide hi Petes Ph oe
SN a
Claws'not strongly and thickly pectimate: :..<22¢ 42.5 222--222 oie sees 34 Claws strongly and thickly pectinate.........---- (380) Ctenochira Forster. (Type, Ctenochira bisinuator Forster. )
Frons with one or two tubercles or horns: J2. 2. J...22S2s2 = eee 35
Frons normal, without a tubercle or horn. Metathorax more or less incompletely areolated; first joint of the flagellum much longer than the second; claws simple, or at most with the
hind claws thinly pectinate toward base. (381) Glypta Gravenhorst.
(Type, Glypta teres Gravenhorst. ) Frons with one tubercle or horn; metathorax more or less areolated with two median carine and two large lateral areas at base; first joint of © flagellum elongate; claws pectinate toward base. (882) Conoblasta Forster. Frons with two tubercles or horns; metathorax exareolate, with only the apical transverse carina present; first joint of flagellum much elongate, nearly as long as joints 2-3 united; claws long, pectinate within. (383) Diplastomorpha Forster.
4
| |
é
No. 1206. ICHNEUMON FLIES-—ASHMEAD. 59
Soe Metathorax not completely areolated ................-.................... 37 Metathorax completely areolated.
Clypeus quite depressed; vertex very narrow; eyes strongly convergent anteriorly; areolet defined but open behind; abdomen with trans- verse impressions only on the first four segments; ventral valve somewhat prominent, but still far from tip of the abdomen.
: (384) Panteles Forster.
Clypeus subconvex; vertex broad; eyes not convergent anteriorly; areolet wholly wanting, the submedian cell longer than the median, the transverse median neryure in hind wings broken far below the middle; abdomen without transverse impressions, closely punctate, QPAGUC HA senses (385) Polysphinctomorpha Ashmead, new genus.
(Type, Polysphinctomorpha luggeri Ashmead, manuscript). 37. Abdomen as in Pimpla, the terminal tergites not prolonged ventrally and not inclosing or hiding the terminal urites _...................---. 38 Abdomen with the terminal tergites prolonged beneath and hiding the terminal urites, or forming a cylinder from which projects the hypopygium that extends far beyond the tip of the abdomen; face not narrowed toward the mouth; eyes not, or only faintly, emarginate within; legs moderately stout, the claws long, pectinate; ovipositor scarcely half the length of the abdomen; body always marked with red. (386) Clistopyga Gravenhorst. 38. Transverse cubital nervure present, the first abscissa of the cubitus forming WioranGuSliMehamelesr’ 22 2 8 ie 2 cule bo se me ah ote nods 39 Transverse cubital nervure wanting, the first branch of the cubitus being inter- stitial with the first abscissa of the radius.
Transverse median neryure in hind wings broken; abdomen with the im- pressions on the segments very feeble or faint.
(387) Acrodactyla Haliday = Oxyrrhexis Forster.
Transverse median nervure in hind wings straight, not broken; abdomen with the impressions on the segments distinct.
(388) Zatypota Forster.
39. Clypeus normal, not projecting into a snout-like ledge anteriorly.........-- 40
Clypeus abnormal, as viewed from the side, projecting forward into a snout- like ledge.
Abdomen subpetiolate, closely opaquely punctate, the segments without distinct, transverse impressions; transverse median nervure in hind wings broken far below the middle.
(389) Zarhynchus Ashmead, new genus. (Type, Tryphon ? nasutus Cresson. ) 40. Face medially tuberculate; mesonotum trilobed; metanotum very short, with an apical transverse carina, slightly interrupted medially, the pos- teriorly face very finely, transversely striate; metanotum and the first four abdominal segments clothed with a dense sericeous UbeScen Cement reas <5 ese (390) Sisyrostolus Kriechbaumer. Face normal; mesonotum not trilobed.
Metanotum with a central longitudinal furrow; abdomen with the trans- verse impressions on the segments well defined, the first segment much longer than wide at apex; last joint of tarsi thickened imme- diately from the base, not longer than the third; basal joint of hind tarsi not longer than the two following joints; onychium large, unusually developed............-- (3891) Polysphincta Grayenhorst.
Metanotum without an areola, at apex trilobed or clavate; abdomen with the transverse impressions not sharply defined, the first segment not longer than wide at apex; last joint of tarsi somewhat thickened
60 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXIII.
at tip only, or in male not at all thickened, longer than the three preceding amen in male longer than the second; onychium not unusually developed .....----------------- (392) Zaglypta Forster,
Tribe Vi XORTDINE
1859-60. Norides, Subfamily, Hotmaren, Kongl. Vets.-Akad. Handl., III, p. 6
1868. Noridoidx, Family 18, Férsrer, Verh. d. naturh. Ver. pr. Rheinl., XXYV, pp. 142 and 168.
1894. Noridini, Tribe IV, Asameap, Proc. Ent. Soc. Wash., ITI, p. 278.
1900. Noridini, Tribe V, AsHMEAD, Smith’s Insects of New Jersey, p. 575.
This tribe is distinguished by the shape of the head, which is quad ‘ate, the temples being broad, and by the peculiar mouth opening formed by the projecting mandibles and the concave or depressed cly- peus, somewhat similar to the mouth opening found in Wesmael’s division Cyclostom? in the family Braconidae.
I have included in the tribe the genera Hehthrus and Nyxeophilus which most authorities place in the subfamily Cryptine, and which seem to form a transition between them and the Pimpline. They are placed here on account of the position of the spiracles of the first abdominal segment, which are placed a¢ or a little before the middle, and not beyond the middle, as in all genuine Cryptines. The inflated front tibiz, too, is a character foqucuily found in this group and rare in the Cryptine.
The group, as a whole, seems to confine itself to attacking the larve of wood-boring Coleoptera. ‘
Twenty-four genera have been placed here, distinguished as follows: _
TABLE OF GENERA.
Areolet in front wings wanting, or small, triangular, or rhomboidal, never large or pentagonal! 228 220. i bees sss Se eee : Areolet in front wings large, pentagonal, or at most subtriangular or subtrapezoidal; anterior tibize in female usually inflated, constricted at base; abdo- men ‘petiolate or subpetiolate: .2-=2--2222.- 2225-22 ee Transverse median nervure in hind wings broken far below the middle; disco cubital nervure in front wings not broken by a stump of a vein.. 4_ Transverse median nervure in hind wings broken at ora little above the middle; disco-cubital nervure broken by a stump of a vein.
Transverse median neryure in front wings interstitial with the basal nervure, the median and submedian cells therefore of an equal length. 220.502 .¢le.S (22-2 Se ee ee
Transverse median nervure in front wings originating before the basa neryure, the submedian cell therefore shorter than the median; metathorax with two transverse carine, the spiracles long oval dorsal carinze of first abdominal segment distinct to near the apex
(393) Nyxeophilus Forster. 3. Metathorax exareolated, at most with only one transverse carina—the apical; | spiracles linear; dorsal carina of first abdominal segment wanting or indicated only at base ..........--- (394) Echthrus Gravenhors 4. Head transverse, the temples narrow -.-/.:222.32 Se ee Head quadrate, the temples broad.
:
No. 1206. ICHNEUMON FLIES—ASHMEAD. 61
Metathorax exareolated, with one transverse carina—the basal; spiracles small, rounded; submedian cell longer than the median, the areolet rather small pentagonal; first abdominal segment short, usually shorter than the second .......-......-- - (395) Holcostizus Forster.
Metathorax with a median area which is, however, confluent with the peti- olar area; submedian cell shorter than the median; first abdominal segment not short, distinctly petiolate.
(896) Cubocephalus Ratzeburg.
5. Metathorax areolated. Seutellum rather flat; anterior tibize deformed, femora incrassated; areolet Wentaconaleyen see sae ee ee (897) Dyseidopus Kriechbaumer. Scutellum gibbous; anterior tibize subinflated, constricted at base; areolet oblique; subrhomboidal.-.----.-- (398) Microtritus Kriechbaumer.
6. Not all the femora short and much swollen, the hind femora always unarmed_ 7 Allthe femora short and much swollen, the hind femora sometimes armed with a tooth beneath. Metathorax areolated; front wings without an areolet; abdomen petiolate, the ovipositor longer than the abdomen. Hind femora armed with a strong tooth beneath; upper hind angles of metathorax toothed or spined- ---- (399) Odontomerus Gravenhorst. Hind femora unarmed but much swollen; hind angles of metathorax normal, not toothed. ---. (400) Anodontomerus Ashmead, new genus. (Type, Aplomerus tibialis Provancher. ) Rema SIT eC sr1ZOr ROTM. sa sec see aeeee ws SEE ake eke tek seek eek 8 Frons with a prominent horn or excrescence. Mesonotum with distinct furrows; metanotum areolated; abdomen petiolate, the ovipositor as long as the abdomen. (401) Ischnoceros Gravenhorst. 8. Front wings without an areolet, the areolet entirely absent. Aibdomengdistmcthysessil ete yaa ce Mesh Oanes Dies SI See eek 12 Abdomen distinctly petiolate. Second recurrent nuryure not angularly broken by a stump of a vein. (402) Clepticus Haliday. (Type, Clepticus pretor Haliday. )
Second recurrent nervure angularly broken by a stump of a vein; stigma scarcely developed; transverse median nervure in hind wings angu- larly broken near the middle; legs long. --- (403) Epixorides Smith.
(Type, Epixorides chalybeator Smith. ) Front wings with an areolet, rarely open behind. Mandibles of an equal length; body stouter and not so elongate .-.--.--- 9 Mandibles of an unequal length; body slender and elongate.
Head not much swollen, subquadrate; metathorax exareolated; abdo- men slender; the ovipositor at the most as longas the abdomen; legs very slender, the hind pair lengthened ~~... (404) . Calliclisis Forster.
9. Clypeus medially lamellate or toothed; metathorax exareolated, or at most with longitudinal carinze, rarely indistinctly areolated. MEeColeh completely closed. jose 4 eee soe eS sesh eee Se eee 10 Areolet open behind. Clypeus anteriorly medially lamellate or toothed; metathorax indis- |
ninety. areolateds cease aaeaees oe See Seas (405) Perosis Forster. a0. Temples posteriorly simple, not tuberculate........------.---.------------ 11 Temples posteriorly tuberculate ....-.-.--- (406) <Achorocephalus Kriechbaumer.
11. Transverse median nervure in hind wings broken below the middle; front tibize moderately thickened but not inflated; middle mesothoracie lobe
3
62 PROCEEDINGS OF THE NATIONAL MUSEUM. VoL. Xxn1T
not projecting above the lateral lobes; petiolar area widely open a the middle; wings often with a brown transverse band.
(407) Xylophrurus Forster.
Transverse median nervure in hind wings broken behind the middle; front tibice
inflated, constricted at base; mesonotum trilobed, the middle lobe—
briefly caniculate; metathorax irregularly arcuately striate.
(408) Gabunia Kriechbaumer. :
12. Metanotum not or very indistinctly areolated; all tibize slender or only slightly thickened; antennz in female without stiff bristles or hairs; abdomi-
nal segments 2 and 3 without transverse impressions; legs slender, —
the posterior pair lengthened: .- 5.22.22 ee eee nee 17
Metanotum usually completely areolated, rarely exareolated; front tibiae much :
thickened or inflated, constricted at base; antennee in female with
rather stiff bristles; abdominal segments 2 and 3 with a more or
less distinct transverse impression.
Metanotum. completely areolated 25-2 22a 2e: Sac ee eee 139
Metanotum) notiareolatedesse=. sau ss ene ee aee = (409) Moansa Tosquinet.
13. Antenne in both sexes short and faintly hairy, the female alone with stiff |
bristles before. apex: Sica ee her ee ee 14 Antenne in both sexes clothed with long shaggy hairs—in male entirely, in
female only toward apex; female antennze ringed with white; ovi- ;| positor longer than the abdomen... ---- (410) Sterotrichus Forster.
14. Transverse median nervure in front wings uniting with the median vein beyond |
the origin of the basal nervure; first abdominal segment without a transverse ridge before apex
first abdominal segment with a transverse ridge before apex. (411) Gonophorus Forster. 15. First abdominal segment medially not emarginate...........-.-----2----.- 16 First abdominal segment medially more or less emarginate. (412) Xylonomus Gravenhorst. i6. Second abdominal segment longer than wide ....---- (413) Merophora Forster. Second abdominal segment not longer than wide.
Head behind the eyes inflated; antennze in both sexes ringed with white; first abdominal segment with two complete carinee; ovipositor as long ‘asthe bedy.2as 2 Sasa ees ee (414) Sichelia Forster.
Head behind the eyes not inflated; antennze not ringed with white; first abdominal segment without complete carine.
(415) Rhadina Forster. 17. Face distinctly narrowed anteriorly; mandibles of an equal length; clypeus at apex strongly impressed; head broadened behind the eyes; abdo-
men more or less sessile, rarely somewhat petiolate. (416) NXorides Gravenhorst. Face not or scarcely narrowed anteriorly; mandibles of an unequal length; clypeus at apex flat, not impressed; head somewhat inflated, slightly narrowed behind the eyes; abdomen narrow, cylindrical, petiolated, with segments 1-5 in female, or 1-7 in male, longer than
WICC. bo. oe oon reees ce 5 Jee Cee (417) Pemenia Holmgren.
; | - :
ete belie ie ee 1 Transverse median nervure in front wings originating before the basal neryure; —
eve
See uae i a ees yO ee N atk TRE A,
ten
| ie
No. 1206. ICHNEUMON FLIES—ASHM EAD. GF
Subfamily DV. LRY PHONIN- A.
1887. Tryphoninx, Subfamily, Cresson, Syn. Hym. North America, p. 47.
1889. Tryphonidx, Family, THomson, Opus. Ent., XIII, pp. 1429 and 1438.
1895. Tryphoninx, Subfamily III, Asumeap, Proce. Ent. Soc. Wash., III, p. 277.
1897. Tryphonine, Subfamily, Davis, Trans. Am. Ent. Soc., XXIV, p. 193.
1900. Tryphonine, Subfamily IV, Asumean, Smith’s Insects of New Jersey, p. 575.
The straight, never elbowed, first abdominal segment, which is usually sessile, and the position of its spiracles, as well as the venation of the front wings, readily distinguish this subfamily from all except the Pimpline andthe Ophioninw. From the former it is easily separated, in the females, by the hidden, or, at most, subexserted, non-prominent ovipositor; from the latter by the much shorter, broader, non-com- pressed abdomen, and a totally different habitus.
Some males, however, are placed with the greatest difficulty, and may be easily confused with those to be found in both the above- mentioned subfamilies.
I know of no good character to easily distinguish them, although the practiced eye, in most cases, is able to place them by comparing them for venational and metathoracic characters peculiar to the females, in the different groups.
The Zryphonine may be divided into ten minor groups or tribes, as follows:
TABLE OF TRIBES.
Basterorupice with one or twovapical spurs: 22 ----. 2-22-22. n2-22- eds oc ee 2 Posterior tibize without apical spurs. Second and third abdominal segments without lunule....Tribe II. Creniscin1. Eeeostenor tibice with only one apical spur:.2:.. 22-022 22.-0ec20 00202015. 4 Posterior tibize with two apical spurs. Abdomen sessile or subsessile, never distinctly petiolate............_.-- 3 Abdomen distinctly petiolate. Claws simple,’not pectinate...----2--..-. 2.2. Tribe I. MEsoLepTint. Glows weCtinate weeescoce . Onc t ei-i5-ccGece Tribe III. Crenopetmint. PERM SEDC CUIMNALCN a Seta se SSS tee ticle Ueto ayed asds Tribe III. Crenoretmint. Claws simple, not pectinate. Mandiblesvbidentaters: 25. Ficlic< ene asc denn i oe Tribe IV. TryPHOoNINI. Nan diblesttridentate ss. ssa hoses tee See Sass cee Tribe V. BaAssInI. Peeninadle tibise-with only one apical ‘spur.-----.--..-.-2-~J..s-2--s22ee-cecee 6 Middle tibize with two apical spurs. HACORO RM aE OS WO GI ams beet Ree Sees SER alles he eR ae 5 Face abnormal, greatly swollen; hind femora usually short and much swollen. Scape lengthened, not short, globose .-.---- Tribe VI. OrrHoceNtTRINI. SCAPO SMOKE ClOWOSeheee eee enter a ee aoa Sse Tribe VII. Exocurnt.
5. Abdomen sessile; dorsum of first and second segments with two parallel carinze. Tribe VIII. Tytecomnrnt.
Abdomen petiolate, the petiole long; dorsum of second segment without carinie; scutellum margined; areolet in front wings not large, subsessile,
Tul Tall. CL cule ever ie eer Sys roe ee Tribe IX. SpHINCTINI.
64 PROCEEDINGS OF THE NATIONAL MUSEUM. VOL. XXIII.
One ace"and: sembtellammaormiell Pa a rs ae Tribe 11. Creniscrnt (part). Face scutatiform; scutellum quadrangular, margined laterally; abdomen long, sessile, the sides parallel or nearly, coarsely punctate; areolet in
front wings usually large, lozengoidal, or diamond-shaped. Tribe X. Meroprint.
Tribed, -MESOLEREINE
1855. Tryphonides homalopi HoLMGREN (part), Kongl. Svensk. Vet.-Akad. Hand1.,_ I, p. 98.
1868. Mesoleptoide, Family 35, Forsrrr, Verh. d. naturh. Ver. pr. Rheinl., XX V, pp. 34 and 197.
1883. Mesoleptina, Tribus, THomson (part), Opus. Ent., LX, pp. 876 and 906.
1894. Mesoleptini, Tribe I, AsHmean, Proc. Ent. Soc. Wash., ITI, p. 277.
1897. Mesoleptini, Tribe, Davis, Trans. Am. Ent. Soc., XXIV, p. 300.
1900. Mesoleptini, Tribe I, AsumEap, Smith’s Insects of New Jersey, p. 575.
In having a distinctly petiolated abdomen this tribe agrees with Ichneumonine, Cryptine, and some in the Ophionine, from all, how- ever, excepting some in the last mentioned, it is at once separated by the straight, no¢ elbowed petiole, and by the position of the spiracles, which are placed a¢ or before the middle, never behind, while from the few genera in the Ophzoninw having the spiracles similarly situated, it is readily distinguished by the non-compressed abdomen, and by the- abdomen in the males not terminating in two long spines.
The only group in the subfamily Z7ryphonine, with which it could be confused, if the other characters made use of in my table are taken into consideration, is the tribe Cfenopelmini, but from this tribe it is separated by the simple, not pectinate, claws.
Thirty-six genera have been recognized, distinguishable by charac-— ters made use of in the following table:
TABLE OF GENERA. Head transverse, the vemples not broad, scarcely half as wide as the width of the CYGS. Sec e cee moet e ee eee ee ee eee eee ee 6
Head quadrate, the temples broad, fully as broad as the width of the eyes. The longer spur of hind tibize as long as or longer than the second joint of
tarsi... no ee ge ett e tee, ee eee 3 The longer spur of hind tibize shorter than the second joint of tarsi. { Front wings with an-areolet..2:.2 <<. 2.) eee 20 Front wings without an areolet............----:- (418) Spanotecnus Forster. — 2. Transverse median nervure in hind wings broken below the middle; metanotum _ incompletely areolated.......-...-.--2.- (419) Eclytus Holmgren. — Transverse median nervure in hind wings broken above the middle; metanotum completely areolated 2.2. 22.222) eee (420) Ichnxops Forster. 3. First recurrent nervure, or the disco-cubital nervure not angularly broken; head — not unusually: swollen-.c:~ te 2 ee ee 4
First recurrent nervure, or the disco-cubital nervure angularly broken; head very much swollen, the yertex posteriorly deeply emarginate.
(421) Polyoncus Forster. —
4. Stigma broad; first joint of the flagellum not longer than the second; metatho-— racic spiracles round, and not lying nearer the external area than —
No. 1206. ICHNEUMON FLIES—ASHMEAD. 65
to the pleural area; first abdominal segment with deep lateral foveze SAMA S ge ete ame ee aie es Seem oie ae Sine loo aec o'ecte ne 2 5 Stigma narrow; first joint of the flagellum distinctly longer than the second; metathoracic spiracles linear and lying nearer the external area than to the pleural area; first abdominal segment usually without deep lateral foveze at base ----.-.......-- (422) Sychnoleter Forster. 5. Transverse median nervure in hind wings broken below the middle.
(423) Laphyroscopus Forster.
Transverse median neryure in hind wings broken at or above the middle. (424) Perilissus Forster.
6. Flagellum 35—40-jointed, usually somewhat thickened; abdomen in female with the last segment so emarginate that the ovipositor lies immediately EDAD The Ui ake Keene fe ee er See ine ee eel oc eer a Flagellum not especially thick; abdomen in female with last segment normal or MOUEIN ALC IN UC Seperate SIR ne keen eee ens Seer oe 8 7. Metanotum at base exareolated; hypopygium in female very prominent. . (425) Polycinetis Forster. Metanotum at base areolated. Front wings without an areolet........2..--------- (426) Homaspis Forster. Front wings with an areolet. Second abdominal segment with two middle carina