Faunitaxys, 9(17), 2021 : 1-6.

Further comments on the elements of the family Palaeoburmesebuthidae Lourenco, 2015 with description of a new species of Spinoburmesebuthus Lourenco, 2017 from Early Cretaceous Burmite amber (Scorpiones)

WILSON R. LOURENCO

Muséum national d’Histoire naturelle, Sorbonne Universités, Institut de Systématique, Evolution, Biodiversité ISYEB), UMR7205- CNRS, MNHN, UPMC, EPHE, CP 53, 57 rue Cuvier, 75005 Paris, France. - wilson.lourenco@mnhn.fr

- ZooBank : http://zoobank.org/58448BD6-79D7-46CE-AFDD-91 EFF2B7D4EF

Keywords:

scorpion; knodelorum; fossil; amber; Early Cretaceous; Myanmar;

Palaeoburmesebuthidae; new species; Spinoburmesebuthus; description.

Abstract. — New comments are proposed on the diversity and morphological variability found within the elements, genera and species, of the Early Cretaceous scorpion family Palaeoburmesebuthidae Lourengo, 2015. A new species belonging to the genus Spinoburmesebuthus Lourengo, 2017 is described rising the total number of species in this family to 13.

Lourencgo W. R., 2021. — Further comments on the elements of the family Palaeoburmesebuthidae Lourengo, 2015 with description of a new species of Spinoburmesebuthus Lourenco, 2017 from Early Cretaceous Burmite amber (Scorpiones). Faunitaxys, 9(17): 1 — 6.

ZooBank: http://zoobank.org/E923F65B-F9FB-4A 8F-97B2-51E74048EES9

Introduction

Among the fossil arthropods found trapped in amber, scorpions can be considered rare. Nevertheless, the very first records can be dated from the early XIX century. New pieces started to become available during the 1980s and in particular the 1990s. These concerned basically scorpions from the Middle and Late Tertiary, from Baltic, Dominican and Mexican amber. Due to the more or less recent age of the amber pieces, the discoveries achieved did not lead to remarkable findings. In fact, many of the scorpions trapped in these types of amber could be easily accommodated in extant groups (for precise details refer to Lourengo, 2009, 2016).

About twenty years ago, the first scorpion trapped in Burmite (Early Cretaceous Burmese amber) became available for study. The specimen was largely incomplete and led only to the creation of a new genus and species, Palaeoburmesebuthus grimaldii Lourenco, 2002 (Lourenco, 2002). This preliminary study was followed by many others since more and more specimens from Burmite started to become available leading to a much better view about this Early Cretaceous scorpion fauna (Lourengo, 2016).

The clarification of the familial status of the genus Palaeoburmesebuthus was only possible several years later after the study of much better preserved specimens. The genus was placed in its own family, Palaeoburmesebuthidae Lourenco, 2015 and a second genus Betaburmesebuthus Louren¢go, 2015 was equally created to accommodate a new lineage of this group (Lourenco, 2015; Lourenco & Beigel, 2015). The number of new species started to progress in an important pace. Indeed in only 16 years 4 species of Palaeoburmesebuthus, 7 of Betaburmesebuthus and a third genus, Spinoburmesebuthus Lourencgo, 2017 was created (see check-list for the species and associated references). The study of several specimens that are almost perfectly preserved, clearly attested their relationship to the buthoids, in particular based on their trichobothrial patterns, which are almost identical to those of several extant buthoids (Lourenco, 2016). Presently the family Palaeoburmesebuthidae is composed of three genera, Palaeoburmesebuthus, Betaburmesebuthus and

Spinoburmesebuthus. A new specimen of Burmite scorpion is presently studied. It leads to the description of a second species belonging to the genus Spinoburmesebuthus. This specimen is remarkably well preserved and brings not only further support to the validity of the genus Spinoburmesebuthus, but also to the relations of the family Palaeoburmesebuthidae to extant lineages.

Materials and methods

The new specimen investigated here is preserved in a rectangular piece of clear yellow amber measuring 35.0 x 25.0 x 17.0 x 5.0 mm. Some organic inclusions and bubbles are present as well as one Acarian element and some micro- Coleopteran. The piece shows a totally complete scorpion and a remarkable number of characters, including the totality (or almost the totality) of trichobothria, are visible in this specimen, including all the bothria, allowing a very precise investigation. Just a few characters, such as the pectines, are less well observable. The schematic drawings provided here are interpretations of what was observable. Illustrations and measurements were produced with the aid of a Wild M5 stereomicroscope equipped with a drawing tube and an ocular micrometer. Measurements follow Stahnke (1970) and are given in mm, and morphological terminology mostly follows Hjelle (1990). Trichobothrial notations follow Vachon (1974). Trichobothria were definitely recorded only when their bothria (areoles) were observed. No supplementary trichobothria was suggested, for example by the presence of transverse hairs.

Comments on the genera and species of the family Palaeoburmesebuthidae

Although the number of contributions to the Burmite scorpions starts to be significant, no previous attention was paid to the possible existence of morphological-type-groups within the genera of the family Palaeoburmesebuthidae. A recent analysis of all described species in the known three genera suggested

2 LOURENCO. — A new Spinoburmesebuthus Lourengo, 2017 (Scorpiones)

the possible existence of morphological-type-groups. Some composed by species with short chelae and telsons and others composed by species with long chelae and telsons. The ratio of chela length vs chela width/depth and telson length vs telson width/depth indicates species distributed over a large range of values (Fig. 1). Naturally, the samples used in the analysis are composed exclusively of the known type material, what is extremely weak. All species are rare and generally only known for a single sex. Nevertheless, the observed existence of possible species-groups seems to be clear. Naturally, all these morphological-type-groups remain totally informal.

Palaeoburmesebuthidae Lourenco, 2015

Genus Palaeoburmesebuthus Lourengo, 2002

— Palaeoburmesebuthus grimaldii Lourengo, 2002

— Palaeoburmesebuthus ohlhoffi Lourengo, 2015a

— Palaeoburmesebuthus longipalpis Lourenco & Rossi, 2017 — Palaeoburmesebuthus knodeli Lourengo, 2018

Genus Betaburmesebuthus Lourenco, 2015 (Lourengo & Beigel, 2015)

— Betaburmesebuthus kobberti Lourengo, 2015 (Lourengo & Beigel, 2015) — Betaburmesebuthus muelleri Lourencgo, 2015b

— Betaburmesebuthus bidentatus Lourenco, 2015b

— Betaburmesebuthus fleissneri Lourengo, 2016 (Lourengo & Velten, 2016a) — Betaburmesebuthus bellus Lourengo, 2016

— Betaburmesebuthus larafleissnerae Lourenco, 2016 (Lourenco & Velten, 2016b)

— Betaburmesebuthus joergi Lourenco & Rossi, 2017

Genus Spinoburmesebuthus Lourengo, 2017 (Lourengo & Velten, 2017)

— Spinoburmesebuthus pohli Lourenco, 2017 (Lourengo & Velten, 2017) — Spinoburmesebuthus knodelorum sp. n.

Systematic description Superfamily Buthoidea C. L. Koch, 1837 Family Palaeoburmesebuthidae Lourenco, 2015

Genus Spinoburmesebuthus Lourengo, 2017

(Fig. 2-5) ZooBank: http://zoobank.org/ECCE73FF-B596-4699-84E3-65414F380F71

Revised diagnosis for the genus Spinoburmesebuthus. — The general morphology is similar to the other two genera of the family Palaeoburmesebuthidae, but also recalls that of some extant buthoid scorpions. The genus Spinoburmesebuthus is however defined on the basis of a combination of characters: A medium to large global size compared to the other elements of the family Palaeoburmesebuthidae with total lengths ranging from 17.62 to 21.76 mm. The two known species are slightly bulk with short pedipalps and metasoma or slender with elongated pedipalps and metasoma; however this can be probably due to sexual dimorphism since the only known specimens are respectively a female and a male. Carapace moderately granular; anterior margin with moderate to strong emargination. Sternum pentagonal in at least one species. Tergites with one median carina and two inconspicuous lateral carinae. Sternites with spiracles ranging from slit-like to oval in one species and semi- oval to almost round in the second species. Metasomal segments I with 10 carinae; segments I] to IV with 8 carinae; segment V with 5 carinae; telson short in female and strongly elongated in male; in all cases with strongly marked spinoid granules and one or two subaculear tooth; setation on metasomal segments moderately to weakly marked. Fixed and movable fingers of pedipalp chela with 9 to 10 series of rounded granules in one species and very sharped granules in the second species; in both cases separated by very conspicuous accessory granules. Trichobothrial pattern shows elements extremely similar to those of extant buthid type A (Vachon 1974): The dorsal trichobothria of femur in beta configuration (Vachon, 1975). For the new species described here almost all

Spinoburmesebuthus knodelorum sp. n.

Spinoburmesebuthus pohli

Betaburmesebuthus joergi Betaburmesebuthus larafleissnerae Betaburmesebuthus bellus Betaburmesebuthus fleissneri Betaburmesebuthus bidentatus Betaburmesebuthus muelleri

Betaburmesebuthus kobberti

Palaeoburmesebuthus knodeli Palaeoburmesebuthus longipalpis Palaeoburmesebuthus ohlhoffi Palaeoburmesebuthus grimaldii

10 4.00

Telson gam Females ge Males

Chela man Females

mums Males

6.00 8.00 Telson or Chela Length/Width

10.00

Fig. 1. Distribution of the species of the family Palaeoburmesebuthidae according to the ratio of length vs width/depth of their chela and telson. See also Table I.

Faunitaxys, 9(17), 2021: 1-6. 3

Fig. 2-5. Spinoburmesebuthus pohli, 2, holotype, type species of the genus (scale bars = 1 mm).

2-4) Trichobothrial pattern. 2) Chela dorso-external aspect. 3) Patella, dorsal aspect. 4) Femur, dorsal aspect. 5) Metasomal segment V and telson, lateral aspect. To notice the strong spinoid granules present in all segments.

Fig. 6-10. Spinoburmesebuthus knodelorum sp. n., 3, holotype (scale bars = 1 mm (6, 8-10); 0.5 mm (7)).

6) Metasomal segments and telson, lateral aspect, showing the spinoid granulations. 7) Chelicera, dorsal aspect. 8-10) Trichobothrial pattern. 8) Chela dorso-external aspect. 9) Patella, dorsal aspect. 10) Femur, dorsal aspect.

4 LOURENCO. — A new Spinoburmesebuthus Lourengo, 2017 (Scorpiones)

trichobothria can be observable; 4 internal, 2 external and five dorsal trichobothria in the femur; 1 internal, 5 dorsal and 7 external trichobothria on patella; no ventral are present; 6 dorso-external and two ventral on chelal hand and 7 on fixed finger can be clearly observed including the internal one. Based on this number it can be suggested that the trichobothrial pattern is orthobothriotaxique. The position of several trichobothria is however quite distinct for the two known species. Tibial spurs present on legs III and IV.

Spinoburmesebuthus knodelorum sp. n.

(Fig. 6-10, 11-16) ZooBank: http://zoobank.org/A286ECF9-BB65-496D-8555-DC8C953D7258

Holotype, 3, probably adult. Included in a rectangular clear block of pale yellow amber that measuring 35.0 x 25.0 x 17.0 x 5.0 mm. Type locality and horizon: Myanmar (Burma), Kachin; precise locality unknown; Lower Cretaceous.

Patronym. — The specific name honors Mr and Mrs Herbert Knodel (Rotonda, USA) who arranged facilities for the study of the type specimen.

Repository. — The type specimen is deposited in the collection of Mr H. Knodel (Rotonda, USA).

Diagnosis. — The new species combines most characters already described for the genus, but can be further diagnosed by the following features: a very slender body and appendages with a total length of 17.62 mm. Marked spinoid granules on femur, patella and chela of pedipalps and dorsal carinae of metasomal segments. Sternum pentagonal. Sternites with small semi-oval to round spiracles. Setation on metasomal segments moderately to strongly marked. Telson granular with two moderately to strongly developted subaculear tubercles. Pectines large with 15-16 teeth. Trichobothrial pattern of buthid type A with a beta configuration (Vachon, 1974, 1975), most certainly orthobothriotaxic. Femur with 5 dorsal 2 external and 4 internal trichobothria; patella with one internal, 5 dorsal and 7 external trichobothria; no supplementary trichobothria are suggested by the observation of fine setae; chela with 8 trichobothria, 6 dorso-external and two ventral; fixed finger with 7 trichobothria, including one internal; trichobothria eb and esb of fixed finger almost at the same level. Tibial spurs present on legs III and IV, strongly marked; tarsi with several thin setae similar to that of some extant buthids such as the genus Ananteris Thorell. Fixed and movable fingers of the pedipalps with sharped granulations recalling diminutive ‘knives’; accessory granulation very strongly sharped, recalling those of Archaeobuthus estephani Lourenco, 2001 from Lebanon Cretaceous amber (Lourenco, 2001).

Description

Coloration. — The scorpion is yellow to slightly reddish-yellow; carapace and tergites yellow to reddish-yellow; metasomal segments reddish to dark reddish; telson reddish with aculeus yellow at the base and reddish at the tip; pedipalps and legs yellow. Ventral aspect dark yellow.

Morphology. — Carapace moderately to strongly granular; anterior margin with a moderately to strongly marked median concavity; posterior edge almost without any emargination. Carinae conspicuous; furrows moderate. Median ocular tubercle clearly anterior to the centre of carapace; median eyes moderate in size and separated by about one ocular diameter. Three pairs of lateral eyes of large size. Sternum pentagonal. Mesosomal tergites moderately to weakly granular, with one median carina; lateral carinae totally inconspicuous; VII with five strongly marked carinae and spinoid granules laterally. Pectines large, with 15-16 teeth; fulcra absent. Sternites

weakly granular with small semi-oval to round spiracles. Metasomal segment I to IV with 10-10-8-8 strongly marked carinae; segment V slender with five carinae; dorsal carinae of segments I-IV with spinoid granules; ventral carina on segment V equally with spinoid granules; dorsal aspect of segments I to V weakly depressed; setation on all segments strongly marked. Curiously, metasomal segment III is shorter than segments I and II (see measurements after the description). Telson with a marked elongated vesicle; moderately granular; aculeus as long as the vesicle and moderately curved; setation strongly marked. Cheliceral dentition only partially visible; fixed and movable fingers with one and two basal tooth; distal teeth moderately long (Vachon, 1963). Pedipalp femur pentacarinate; patella with 6-7 carinae; internal face of femur and patella with strongly marked spinoid granules. Chela with moderately marked carinae; all faces weakly granular; internal face with 2-3 spinoid granules. Fixed and movable fingers each with one series, divided in 9-10 sub-series, of small knife-shaped granules separated by conspicuous knife-shaped accessory granules; extremity of fingers with stronger knife- shaped granules; setation of pedipalps moderately marked. Trichobothriotaxy of type A (Vachon, 1974) defined for extants buthids; dorsal trichobothria disposed in beta (B) configuration (Vachon, 1975). General pattern most certainly orthobothriotaxic. Femur with 5 dorsal 2 external and 4 internal trichobothria; patella with one internal, 5 dorsal and 7 external trichobothria; no supplementary trichobothria are suggested by the observation of fine setae; chela with 8 trichobothria, 6 dorso-external and two ventral; fixed finger with 7 trichobothria, including one internal; trichobothria eb and esb of fixed finger almost at the same level. Tibial spurs present on legs III and IV, strongly marked; tarsi with several thin setae.

Morphometric values (mm), holotype.

— Total length: 17.62 (including telson). — Carapace: length 2.24, anterior width 1.07, posterior width 2.17. — Mesosoma: length 4.54. — Metasomal segments I: length 1.37, depth 0.67; II: length 1.50, depth 0.54; II: length 1.14, depth 0.57; IV: length 1.71, depth 0.60; V: length 2.57, depth 0.60. ~ Telson: length 2.55. — Vesicle: depth 0.37. —Pedipalp femur length 1.94, width 0.47; patella length 2.10, width 0.57; chela length 3.74, width 0.37; — Movable finger: length 2.95.

Acknowledgements

I am most grateful to Mr H. Knodel (Rotonda, USA) for arranging facilities for the study of the specimen, and for the preparation of the photos and to Lucienne Wilmé (Missouri Botanical Garden, USA) for the preparation of the graph (Fig. 1).

References

Hjelle J. T., 1990. — Anatomy and morphology (p. 9-63). Jn: Polis G. A. (ed.), The Biology of Scorpions. Stanford: Stanford University Press, 587 p.

Lourengo W. R., 2001. — A remarkable scorpion fossil from Lebanon amber. Implications for the phylogeny of Buthoidea. Comptes Rendus de 1l’Académie des Sciences, Paris, Sciences de la Terre et des planétes, 332: 641-646.

Faunitaxys, 9(17), 2021 : 1-6. 5

4 ig te is eS oo bi ee Ps Ee ar | : Fig. 11-16. Spinoburmesebuthus knodelorum sp. n., 3, holotype.

11) Habitus, dorsal aspect. 12) Ventral aspect, showing pectines. 13) Metasomal segments and telson, lateral aspect. 14) Segment V and telson in detail. 15) Left Chela,

showing the spinoid granulations on fixed and movable fingers. 16) Extremity of the fingers, in detail, showing the conspicuous knife-shaped accessory granules.

Lourenco W. R., 2002. — The first scorpion fossil from the Cretaceous amber of Myanmar (Burma). New implications for the phylogeny of Buthoidea. Comptes Rendus Palevol (Académie des Sciences) Paris, 1: 97-101.

Lourenco W. R., 2009. — A synopsis of the amber scorpions, with special reference to the Baltic fauna. In: Zugleich Kataloge der oberésterreichischen Landermuseen. Denisia, 26 (n. ser.), 86: 131-136.

Lourenco W. R., 2015a. — Clarification of the familial status of the genus Palaeoburmesebuthus Lourengo, 2002 from Cretaceous Burmese amber (Scorpiones: Archaeobuthidae: Palaeoburmesebuthinae). Beitrdge zur Araneologie, 9: 465-475.

Lourengo W. R., 2015b. — A new contribution to the knowledge of Cretaceous Burmese amber scorpions with the description of two new species of Betaburmesebuthus Lourengo, 2015 (Scorpiones: Archaeobuthidae: Palaeoburmesebuthinae). Arachnida — Rivista Aracnologica Italiana, \(3): 27-36.

Lourenco W. R., 2016. —A preliminary synopsis on amber scorpions with special reference to Burmite species: an extraordinary development of our knowledge in only 20 years. Zookeys, 600: 75-87.

Lourenco W. R., 2018. — A further new species of Palaeoburmesebuthus Lourencgo, 2002 (Scorpiones: Palaeoburmesebuthidae) from Burmite. Revista Ibérica de Aracnologia, 32: 51-54.

6 LOURENCO. — A new Spinoburmesebuthus Lourengo, 2017 (Scorpiones)

Lourenco W. R. & Beigel A., 2015. —A new genus and species of Palaeoburmesebuthinae Lourenco, 2014 (Scorpiones: Archaeobuthidae:) from Cretaceous amber of Myanmar. Beitrdge zur Araneologie, 9: 476-480.

Lourenco W. R. & Rossi A., 2017. — Two more new species of Burmese amber scorpions of the family Palaeoburmesebuthidae Lourengo, 2015 (Scorpiones). Arachnida - Rivista Aracnologica Italiana, 13: 11-21.

Lourengo W. R. & Velten J., 2016a. — One more new species of Betaburmesebuthus Lourencgo, 2015 (Scorpiones: Palaeoburmesebuthinae) from Cretaceous burmite. Arachnida — Rivista Aracnologica Italiana, 6: 4-11.

Lourenco W. R. & Velten J., 2016b. — A sixth new species of Cretaceous Burmese amber scorpion, genus Betaburmesebuthus Lourengo, 2015 (Scorpiones: Palaeoburmesebuthidae). Arachnida — Rivista Aracnologica Italiana, 10: 10-17.

Lourenco W. R. & Velten J., 2017.— One more new genus and species of fossil scorpion from Burmese Cretaceous amber

belonging to the family Palaeoburmesebuthidae. Arachnida — Rivista Aracnologica Italiana, 13: 2-10.

Stahnke H. L., 1970.— Scorpion nomenclature and mensuration. Entomological News, 81: 297-316.

Vachon M., 1963. — De Vutilité, en systématique, d’une nomenclature des dents des chélicéres chez les Scorpions. Bulletin du Muséum national d’Histoire naturelle, Paris, 2e sér., 35 (2) : 161-166.

Vachon M., 1974. — Etude des caractéres utilisés pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Muséum national d Histoire naturelle, Paris, 3e sér., n° 140, Zool. 104 : 857-958.

Vachon M., 1975. — Sur l'utilisation de la trichobothriotaxie du bras des pédipalpes des Scorpions (Arachnides) dans le classement des genres de la famille des Buthidae Simon. Comptes Rendus des Séances de l’Académie des Sciences, 281 (D): 1597-1599.

Table I. Ratios (R) between Telson length (L) and width or depth (W) and Chela length (L) and width or depth (W)

Palaeoburmesebuthus grimaldii Lourenco, 2002 Telson — L= 1.45, W = 0.29 — R= 5.00

Palaeoburmesebuthus ohlhoffi Lourenco, 2015 Telson — L= 3.16, W =0.34-R=9.29 Chela — L = 2.26, W = 0.34-— R = 6.65

Palaeoburmesebuthus longipalpis Lourengo & Rossi, 2017 Chela — L = 2.42, W=0.19—R=12.74

Palaeoburmesebuthus knodeli Lourengo, 2018 Telson — L= 1.67, W = 0.40 — R= 4.18 Chela — L= 2.81, W = 0.47 -—R =5.98

Betaburmesebuthus kobberti Lourengo, 2015 Telson — L= 1.47, W = 0.30 —-R = 4.90 Chela — L= 2.14, W = 0.34— R = 6.30

Betaburmesebuthus muelleri Lourenco, 2015 Telson — L= 1.34, W = 0.34-—R =3.94 Chela — L= 1.74, W=0.34-—R=5.12

Betaburmesebuthus bidentatus Lourengo, 2015 Chela — L = 3.47, W = 0.47 —-R=7.38

Betaburmesebuthus fleissneri Lourenco, 2016 Telson — L= 1.62, W=0.34-—R=4.76 Chela — L = 2.32, W =0.34 —-R = 6.82

Betaburmesebuthus bellus Lourenco, 2016 Telson —- L= 1.47, W=0.37 -R=3.97 Chela — L = 2.24, W =0.34-R=6.59

Betaburmesebuthus larafleissnerae Lourengo, 2016 Telson — L= 1.67, W=0.40-R=4.17 Chela — L = 2.27, W =0.40 —-R =5.68

Betaburmesebuthus joergi Lourenco & Rossi, 2017 Telson — L= 2.54, W = 0.62 -R=4.10 Chela — L = 3.60, W = 0.74 —-R = 4.86

Spinoburmesebuthus pohli Lourenco, 2017 Telson — L= 2.27, W =0.70 —-R=3.24 Chela — L = 3.74, W =0.54 —R = 6.93

Spinoburmesebuthus knodelorum sp. n. Telson — L= 2.55, W=0.37-R=6.90 Chela — L = 3.74, W =0.37 —-R = 10.10

Résumé

Lourengo W. R., 2021. — Considérations supplémentaires sur les éléments de la famille des Palaeoburmesebuthidae Lourenco, 2015 et description d’une nouvelle espéce de Spinoburmesebuthus Lourengo, 2017 de ’ambre du Crétacé inférieur de la Birmanie (Scorpiones). Faunitaxys, 9(17) : 1-6.

Des nouvelles considérations sont proposées sur la diversité et la variabilite morphologique retrouvée chez les éléments, genres et espéeces, de la famille Palaeoburmesebuthidae Lourenco, 2015, du Crétacé inférieur de la Birmanie. Une nouvelle espéce appartenant au genre Spinoburmesebuthus Lourengo, 2017 est décrite, élevant ainsi le nombre total d’espéces connues a 13.

Mots-clés. — Scorpion, fossile, Crétacé inférieur, Palaecoburmesebuthidae, Spinoburmesebuthus, knodelorum, ambre, Myanmar, nouvelle espéce,

description.

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Lourengo W. R., 2021.—Une nouvelle espéce appartenant au genre Buthus Leach, 1815 (Scorpiones : Buthidae) collectée dans le Parc Naturel de la ‘Serra da Estrela’ au Centre du Portugal. Faunitaxys, 9(13) : 1-7.

Lourenco W. R. & Velten J., 2021.— One more new genus and species of scorpion from Early Cretaceous Burmese amber (Scorpiones: Protoischnuridae). Faunitaxys, 9(14): 1—5.

Théry T., 2021.—Description of Eucurtiopsis marysae n. sp., a singular species of Chlamydopsinae from the Philippines (Coleoptera, Histeridae). Faunitaxys, 9(15): 1 —5.

Cumming R. T. & Le Tirant S., 2021. — Review of the Cretaceous tArchaeatropidae and +Empheriidae and description of a new genus and species from Burmese amber (Psocoptera). Faunitaxys, 9(16): 1 — 11.

Faunitaxys est échangée avec les revues suivantes (« print versions ») :

— Annali del Museo Civico di Storia Naturale G. Doria (Italie)

— Boletin de la Asociacion espafiola de Entomologia (Espagne)

— Boletin de la Sociedad Andaluza de Entomologia (Espagne)

— Bollettino del Museo di Storia Naturale di Venezia (Italie)

— Bulletin de la Société linnéenne de Lyon (France)

— Bulletin of Insectology (Italie)

— Heteropterus Rev. Entomol. (Espagne)

— Israel Journal of Entomology (Israel)

— Klapalekiana (République Tchéque)

— Koleopterologische Rundschau (Allemagne)

— Memorie del Museo Civico di Storia Naturale di Verona (Italie) — Nova Supplementa Entomologica (Allemagne)

— Proceedings of the Entomological Society of Washington (USA) — Revue suisse de Zoologie (Suisse)

— Spixiana (Allemagne)

— Stuttgarter Beitrage zur Naturkunde A, Biologie (Allemagne)

— Zoosystematica Rossica (Russie)

Faunitaxys

Volume 9, Numero 17, Mai 2021

SOMMAIRE

Considérations supplémentaires sur les éléments de la famille des Palaeoburmesebuthidae Lourengo, 2015 et description d’une nouvelle espéce de Spinoburmesebuthus Lourencgo, 2017 de l’ambre du Crétacé inférieur de la Birmanie (Scorpiones).

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CONTENTS

Further comments on the elements of the family Palaeoburmesebuthidae Lourenco, 2015 with description of a new species of Spinoburmesebuthus Lourengo, 2017 from Early Cretaceous Burmite amber (Scorpiones).

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Illustration de la couverture : Spinoburmesebuthus knodelorum Lourengo sp. n. dans l’ambre du Crétacé inférieur de la Birmanie.

Publié par |’ Association Frangaise de Cartographie de la Faune et de la Flore (AFCFF)